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Fitz Alumni

PhD Graduates

2010 - present

Barquete Costa, Viviane. Dec 2012. Using stable isotopes as a tool to understand the trophic relationships and movements of seabirds off southern Africa. (Supervisors: Peter Ryan and Ross Wanless).

Baum, Julia. Dec 2016. The influence of location on the structure and functioning of private land conservation networks in the Western Cape Province of South Africa. (Supervisor: Graeme Cumming).

Braby, Justine. June 2011. The biology and conservation of the Damara Tern Sterna balaenerum in Namibia. (Supervisors: Les Underhill and Rob Simmons).

Chaskda, Adams. Dec 2011. Determinants and consequences of territory quality in the Bar-throated Apalis Apalis thoracica. (Supervisors: Penn Lloyd and Phil Hockey).

Clements, Hayley. Dec 2016. Multi-scale, social-ecological influences on private land conservation in South Africa. (Supervisor: Graeme Cumming).

Davies, Owen. June 2015. Taxonomy, phylogeny and biogeorgraphy of cisticolas (Cisticola spp.). (Supervisors: Tim Crowe and Rauri Bowie).

Cohen, Callan. June 2011. The phylogenetics, taxonomy and biogeography of  African arid zone terrestrial birds: the bustards (Otididae), sandgrouse (Pteroclidae), coursers (Glareolidae) and Stone Partridge (Ptilopachus). (Supervisors: Tim Crowe and Rauri Bowie).

de Ponte Machado, Marta. June 2010. Population dynamics of Great White Pelicans on in Southern Africa - causative factors and influence on other seabirds. (Supervisors: Les Underhill, Peter Ryan, Rob Crawford and Rauri Bowie).

Garcia-Heras, Marie-Sophie. July 2017. Environmental factors influencing the breeding and health of a predator endemic to southern Africa: the endangered Black Harrier Circus maurus. (Supervisors: Rob Simmons, Arjun Amar, Beatriz Arroyo and Francois Mougeot).

Henry, Dominic. June 2016. Waterbirds and nutrient dispersal in isiMangaliso Wetland Park. (Supervisor: Graeme Cumming).

Joseph, Grant. Dec 2012. Understanding pattern-process relationships in a heterogeneous landscape: effects of termitaria on diversity and disturbance regimes in miombo woodlands of northern Zimbabwe. (Supervisor: Graeme Cumming).

Jones, Genevieve. Dec 2011. Individual variation in reproductive success in the Wandering Albatross. (Supervisor: Peter Ryan).

Kaliba, Potiphar. June 2014. Faunal turnover between east and southern African terrestrial vertebrates: is Malawi the geographical break? (Supervisors: Rauri Bowie and Tim Crowe).

Krüger, Sonja. Dec 2014. An investigation into the decline of the Bearded Vulture Gypaetus barbatus in Southern Africa. (Supervisors: Arjun Amar and Rob Simmons).

Little, Ian. Dec 2011. Bird reproductive success and faunal habitat selection as tools for under-standing the impacts of land-use management on moist highland grassland biodiversity in South Africa. (Supervisors: Phil Hockey and Ray Jansen).

Mandiwana-Neudani, Tshifhiwa. June 2014. Taxonomy, phylogeny and biogeography of francolins (‘Francolinus’ spp.) Aves: Order Galliformes, Family: Phasianidae. (Supervisors: Tim Crowe and Rauri Bowie).

McInnes, Alistair. Dec 2016. Fine-scale drivers of African Penguin prey dynamics in Algoa Bay, South Africa, and their impacts on penguin foraging ecology. (Supervisors: Lorien Pichegru and Peter Ryan).

Murgatroyd, Megan. June 2016. Ecology of the Verreaux's Eagle in natural and agriculturally transformed habitats in South Africa.  (Supervisors: Les Underhill and Arjun Amar).

Ndlovu, Mduduzi. Dec 2012. Environmental influences on moult and movement strategies in southern African waterfowl. (Supervisor: Graeme Cumming).

Nelson-Flower, Martha. June 2010. Kinship and its consequences in the cooperatively breeding Southern Pied Babbler Turdoides bicolor (Supervisors: Phil Hockey and Mandy Ridley).

Nupen, Lisa. Dec 2014.  Comparative conservation genetics and evolutionary history of threatened, endemic southern African seabirds in the Benguela Current Upwelling Ecosystem: range-wide phylogeography, gene flow and population genetics based on nuclear and mitochondrial DNA. (Supervisors: Peter Ryan, Rauri Bowie and Jacqui Bishop).

Oatley, Graeme. Dec 2011. Taxonomy, phylogeny and eco-biogeography of southern African white-eyes (Zosterops spp.) Aves: Order Passeriformes family: Zosteropidae. (Supervisors: Tim Crowe and Rauri Bowie).

Okanga, Sharon. Dec 2013. The influences of host community, urbanization and water quality on avian malaria ecology in South African passerines (Supervisors: Graeme Cumming and Phil Hockey).

Reynolds, Chevonne. June 2016. Understanding the relationship between waterfowl movement and dispersal of freshwater aquatic organisms in southern Africa (Supervisor: Graeme Cumming)

Ribeiro, Angela. Dec 2011.  Ecological and evolutionary processes in two southern African endemic birds. (Supervisors: Rauri Bowie, Penn Lloyd and Phil Hockey).

Rollinson, Dominic. July 2017. Understanding and mitigating seabird bycatch in the South African pelagic longline fishery. (Supervisors: Peter Ryan and Ross Wanless)

Shaw, Jessica. June 2013.  Powerlines in the Karoo: conserving Ludwig's Bustard.  (Supervisors: Peter Ryan and Andrew Jenkins).

Tate, Gareth. Dec 2016. Exploring the maintenance of plumage polymorphism in the Black Sparrowhawk Accipiter melanoleucus. (Supervisors: Arjun Amar and Jacqui Bishop).

Thompson, Alexander. Dec 2013. Maternal investment and its effects on parent-offspring conflict in the cooperatively breeding Pied Babbler (Turdoides bicolor) (Supervisors: Peter Ryan, Phil Hockey and Nikki Raihani).

van Eeden, Rowen. July 2017. Understanding the decline of Martial Eagles Polemaetus bellicosus in the  Kruger National Park, South Africa. (Supervisor: Arjun Amar).


Barnes, Keith. 2007. The evolution of Africa's larks Alaudidae. (Supervisor: Peter Ryan).

Bowie, Rauri. 2003. Birds, molecules and evolutionary patterns among Africa's islands In the sky. (Supervisor: Tim Crowe).

Covas, Rita. 2002. Life histories and sociality in the Sociable Weaver. (Supervisor: Morné du Plessis).

Jansen, Raymond. 2001. Population Biology, Behavioural Ecology and Management of the Redwing Francolin Francolinus levaillantii and Swainson's Spurfowl Pternistis swainsonii.(Supervisor: Tim Crowe).

Loewenthal, Doug. 2007. Population dynamics and conservation of the African Black Oystercatcher Haematopus moquini. (Supervisor: Phil Hockey).

Muchai, Muchane. 2002. Going through the motions: The impacts of frequent fires and grazing pressure on reproduction by montane grassland birds. (Supervisor: Morné du Plessis).

Nel, Deon. 2002. The impact of longline fishing on the seabirds breeding on Marion Island. (Supervisor: Peter Ryan).

Petersen, Samantha. 2008. Assessing and managing the impacts of long-line fishing on seabirds, turtles and sharks off southern Africa. (Supervisors: Peter Ryan, Les Underhill, Ronel and Len Compagno).

Prendini, Lorenzo. 2002. The taxonomy, phylogenetics, biogeography and functional morphology of the Southern African burrowing scorpions, Genus Opistophthalmus C.l. Koch (Scorpiones: Scorpionidae). (Supervisor: Tim Crowe).

Seymour, Colleen. 2006. The role of Acacia erioloba in preserving biodiversity in the Kalahari, Southern Africa. (Supervisor: Richard Dean).

Techow, Mareille. 2007. Phylogeny and phylogeography of four Southern Ocean petrels. (Supervisors: Peter Ryan & Colleen O'Ryan).

van Zyl, Hugo. 2007. Property price approaches to the valuation of urban wetlands: theoretical implications and property considerations. (Supervisor: Jane Turpie).

Wanless, Ross. 2007. The impacts of introduced mice on birds at Gough Island. (Supervisor: Peter Ryan).


Adams, Nigel. 1990. Feeding biology and energetics of King Aptenodytes patagonicus and Gentoo Pygoscelis papua Penguins at sub-Antarctic Marion Island. (Supervisors: Roy Siegfried, John Cooper and Robert Prys-Jones).

Dean, Richard. 1995. Where birds are rare or fill the air: The protection of the endemic and the nomadic avifaunas of the Karoo. (Supervisor: Roy Siegfried).

de Klerk, Helen. 1999. Biogeography and conservation of terrestrial Afrotropical birds. (Supervisor: Tim Crowe).

du Plessis, Morné.  1990. Behavioural ecology of the Redbilled Woodhoopoe Phoeniculus purpureus in South Africa. (Supervisor: Phil Hockey).

Fabricius, Christo. 1997. The impact of land use of biodiversity In Xeric Succulent Thicket, South Africa. (Supervisor: Phil Hockey).

Jackson, Susan. 1990. Seabird digestive physiology in relation to foraging ecology. (Supervisors: Roy Siegfried and David Duffy).

Jenkins, Andrew. 1998. Behavioural ecology of Peregrine and Lanner Falcons in South Africa. (Supervisors: Roy Siegfried, Phil Hockey and Alan Kemp).

Kaletja, Bozena. 1992. Aspects of the ecology of migrant shorebirds (Aves: Charadrii) at the Berg River estuary, South Africa. (Supervisor: Phil Hockey).

Little, Rob. 1992. Population genetics, behavioural ecology and management of the Greywing Francolin Francolinus africanus.  (Supervisor: Tim Crowe).

Lloyd, Penn. 1998. A study of the ecology of the Namaqua Sandgrouse and other arid-zone birds. (Supervisor: Tim Crowe).

Malan, Gerard. 1995. Cooperative breeding and delayed dispersal in the Pale Chanting Goshawk Melierax canorus. (Supervisors: Tim Crowe and Rob Little).

Macdonald, Ian. 1991. Conservation implications of the invasion of southern Africa by alien organisms. (Supervisor: Roy Siegfried).

Milton, Sue. 1993. Studies of herbivory and vegetation change in Karoo shrublands. (Supervisor: Roy Siegfried).

Rebelo, Tony. 1992. Preservation of the Cape Flora: status, causes of rarity, ideals and priorities. (Supervisor: Roy Siegfried).

Ryan, Peter. 1992. The ecology and evolution of Nesospiza Buntings. (Supervisors: Roy Siegfried and John Cooper).

Velazquez, Claudio. 1993. Ecology and management of waterbirds at commercial saltpans in South Africa. (Supervisor: Phil Hockey).

Turpie, Jane. 1994. Comparative foraging ecology of two broad ranging migrants, Grey Plover Pluvialis squatarola and Whimbrel Numenius phaeopus (Aves: Charadrii) in tropical and temperate latitudes of the western Indian Ocean. (Supervisor: Phil Hockey).


Abrams, Ronald. 1985. The structure of pelagic seabird assemblages in the African sector of the Southern Ocean. (Supervisor: Roy Siegfried).

Bosman, Alison. 1987. Avian determinants of rocky intertidal community structure in South Africa. (Supervisor: Phil Hockey).

Brown, Chris. 1988. Ecological energetics of Eudyptes Penguins at Marion Island. (Supervisors: Roy Siegfried and John Cooper).

Burger, Alan. 1980. Behavioural ecology of Lesser Sheathbills Chionis minor at Marion Island. (Supervisor: Roy Siegfried).

Guillet, Alfredo. 1986. Biogeography and ecology of African waterbirds. (Supervisor: Tim Crowe).

Hockey, Phil. 1983. Ecology of the African Black oystercatcher Haematopus moquini. (Supervisor: Roy Siegfried).

Knight, Richard. 1988. Aspects of plant dispersal in the southwestern Cape with particular reference to the roles of birds as dispersal agents. (Supervisor: Roy Siegfried).

Williams, Tony. 1980. Breeding biology of Eudyptes penguins with particular reference to egg-size dimorphism. (Supervisor: Roy Siegfried).

Wilson, Rory. 1986. Breeding Jackass Penguins Spheniscus demersus as pelagic predators.(Supervisor: Roy Siegfried).


Crowe, Tim. 1978. Evolution and ecology of guineafowl. (Supervisor: Roy Siegfried).

Elliott, Clive. 1973. Biology of the Cape Weaver Ploceus capensis with special reference to its polygynous mating system. (Supervisor: Roy Siegfried).

Farkas, Tibor. 1975. Comparative study of avian communities inhabiting natural and artificial habitats in the eastern part of the Orange river project area. (Supervisor: Roy Siegfried).

Jarvis, Mike. 1971. Ethology and ecology of the South African Gannet Sula capensis. (Supervisor: Roy Siegfried).

Puttick, Gillian. 1979. Feeding ecology of Curlew Sandpipers Calidris ferruginea in the southwestern Cape. (Supervisor: Roy Siegfried).


Siegfried, Roy. 1969. Ecology of the Cattle Egret Ardeola ibis with special reference to population dynamics. (Supervisor: Gerry Broekhuysen).