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Director

Prof Peter Ryan

Prof. Peter G. Ryan
BSc (Hons), MSc, PhD (Cape Town)

John Day Building: 2.03.1
Tel:  +27 (0)21 650 2966
Fax: +27 (0)21 650 3295

Email: peter.ryan@uct.ac.za

Activities and research interests

General ornithology, seabird-fishery interactions, evolutionary ecology (focusing on resolving closely-related species complexes), marine debris, solid waste management, the biology of oceanic islands.

Peter Ryan graduated from the University of Cape Town, researching the impacts of ingested plastic on seabirds (MSc) and the evolutionary ecology of buntings in the Tristan archipelago (PhD). After a post-doc at the University of California he was appointed academic Coordinator of the Masters Programme in Conservation Biology at the Fitztitute in 1993. Although he mostly works on seabirds and their conservation, he has wide-ranging interests in avian biology, and still maintains an interest in plastic pollution, especially in marine systems. He has served as the President of BirdLife South Africa and is an associate editor of Antarctic Science and Bird Conservation International. He has been an honorary Conservation Officer at Tristan da Cunha since 1989, and is a member of Tristan’s Biodiversity Advisory Group. Peter has spent more than a year on Inaccessible Island, studying the island's endemic buntings as well as conducting surveys of threatened birds and conducting control programmes against alien plants. He is a keen birder and believes in the importance of promoting amateur involvement in ornithology. He has written several books about birds and their identification, and is scientific advisor to African Birdlife, a popular magazine about birds produced by BirdLife South Africa.

Research programmes

Island conservation , Seabird conservation , Systematics and biogeography , Rarity & conservation of African birds , Co-operative breeding and sociality in birds

Current students

Doctoral

Kate Carstens: Breeding dynamics, dispersal and fine-scale genetic structure in the endangered Southern Ground-Hornbill  Bucorvus leadbeateri .  (Co-supervisor: Alan Kemp)

Ben Dilley: Breeding success and abundance of burrow-nesting petrels at Marion Island.

Selena Flores: The effects of anthropogenic disturbance of beaches on breeding shorebirds in the high-tourism Garden Route region of South Africa. (Co-supervisor: Mark Brown)

Davide Gaglio: Link between population dynamics, foraging behaviour and food abundance and distribution in the Swift Tern Thalasseus bergii (Co-supervisors: Timothée Cook, Richard Sherley).

Dominic Rollinson : Reducing seabird bycatch from the longline fishing industry (Co-supervisor: Ross Wanless)

Kim Stevens: The foraging ecology and breeding success of Grey-Headed albatross  Thalassarche chrysostoma  on Marion Island.

Tanja van de Ven: Implications of climate change for the reproductive behaviour and performance of the desert-dwelling Southern Yellow-billed Hornbill (Tockus leucomelas) (Co-supervisors: Susie Cunningham, Andrew McKechnie).

Otto Whitehead: Foraging ecology and diving behaviour of crested penguins at Marion Island (Co-supervisor: Yan Ropert-Coudert).

Masters

Andrew de Blocq van Scheltinga: Assessing the impacts of boat-based tourism on waterbirds at De Hoop Vlei. (Co-supervisor: Robert Thomson)

Ditiro Moloto: Does the structure of flight feathers differ among Procellariformes that fly underwater?

Laurie Johnson: Assessing the effect of feather wear on the carbon and nitrogen isotope values, and the use of the stable isotope technique to determine the structure of the food-web in the Namibian Island's Marine Protected Area.

Assessing the effect of feather wear on the carbon and nitrogen isotope values, and the use of the stable isotope technique to determine the structure of the food-web in the Namibian Island’s Marine Protected Area - See more at: http://www.fitzpatrick.uct.ac.za/fitz/students/masters/johnson#sthash.QBr2PxMB.dpuf
Assessing the effect of feather wear on the carbon and nitrogen isotope values, and the use of the stable isotope technique to determine the structure of the food-web in the Namibian Island’s Marine Protected Area - See more at: http://www.fitzpatrick.uct.ac.za/fitz/students/masters/johnson#sthash.QBr2PxMB.dpuf
Assessing the effect of feather wear on the carbon and nitrogen isotope values, and the use of the stable isotope technique to determine the structure of the food-web in the Namibian Island’s Marine Protected Area - See more at: http://www.fitzpatrick.uct.ac.za/fitz/students/masters/johnson#sthash.QBr2PxMB.dpuf

John Pallett: Collisions of large birds, especially bustards, with power lines in Namibia: significance and possible solutions (Co-supervisor: Ann Scott).

Noëlle Tubbs: Heat tolerance in the African Penguin (Spheniscus demersus) in the face of climate change (Co-supervisors: Lorien Pichegru, Jonathan Green).

Conservation Biology Masters

Tendai Chinho: Testing the information centre hypothesis in a colonial seabird: foraging information transfer in breeding Cape gannet pairs: (Co-supervisor: Lorien Pichegru)

Corey Jeal: Testing the information centre hypothesis in a colonial seabird: foraging information transfer in breeding Cape gannet pairs: (Co-supervisor: Samantha Ralston-Paton)

Matthew Macray: Electric fence-induced mortality of leopard tortoises (Stigmochelys pardalis) in Baviaanskloof, South Africa (Co-supervisor: Samantha Ralston-Paton)

esting the information centre hypothesis in a colonial seabird: foraging information transfer in breeding Cape gannet pairs. Supervisor - See more at: http://www.fitzpatrick.uct.ac.za/fitz/students/consebio2017/chinho#sthash.c1yXrayt.dpuf

Peer-reviewed publications

Plastic pollution (37 papers)

Fazey, F. and Ryan, P.G. 2016. Debris size and buoyancy influence the dispersal distance of stranded litter. Marine Pollution Bulletin 110:371-377.

Fazey, F. and Ryan, P.G. 2016. Biofouling on buoyant marine plastics: an experimental study into the effect of size on surface longevity. Environmental Pollution 210: 354-360.

Ryan, P.G., Cole, G., Spiby, K., Nel, R., Osborne, A. and Perold, V. 2016. Impacts of plastic ingestion on post-hatchling loggerhead turtles off South Africa. Marine Pollution Bulletin 107: 155-160.

Ryan, P.G., de Bruyn, P.J.N. and Bester, M.N. 2016. Regional differences in plastic ingestion among Southern Ocean fur seals and albatrosses. Marine Pollution Bulletin 104: 207-210.

Witteveen, M., Brown, M. and Ryan, P.G. 2016. Anthropogenic debris in the nests of Kelp Gulls in South Africa. Marine Pollution Bulletin doi: 10.1016/j.marpolbul.2016.10.052.

Ryan, P.G. 2015. Does size and buoyancy affect the long-distance transport of floating debris? Environmental Research Letters 10: 084019 (doi:10.1088/1748-9326/10/8/084019)

Ryan, P.G. 2015. How quickly do albatrosses and petrels digest plastic particles? Environmental Pollution (doi: 10.1016/j.envpol.2015.08.005)

Ryan, P.G. 2014. Litter survey detects the South Atlantic ‘garbage patch’. Marine Pollution Bulletin 79: 220-224.

Ryan, P.G., Lamprecht, A., Swanepoel, D. & Moloney, C.L. 2014. The effect of fine-scale sampling frequency on estimates of beach litter accumulation. Marine Pollution Bulletin 88: 249-254.

Ryan, P.G., Musker, S. & Rink, A. 2014. Low densities of drifting litter in the African sector of the Southern Ocean. Marine Pollution Bulletin 89: 16-19.

Eriksen, M., Lebreton, L.C.M., Carson, H.S., Thiel, M., Moore, C.J., Borerro, J.C., Galgani, F., Ryan, P.G. & Reissner, J. 2014. Plastic pollution in the world’s oceans: more than 5 trillion plastic pieces weighing over 250,000 tons afloat at sea. PLoS ONE 9(12): e111913.

Vegter, A.C., Barletta, M., Beck, C., Borrero, J., Burton, H., Campbell, M.L., Eriksen, M., Eriksson, C., Estrades, A., Gilardi, K.V.K., Hardesty, B.D., Ivar do Sul, J.A., Lavers, J.L., Lazar, B., Lebreton, L., Nichols, W.J., Ribic, C.A., Ryan, P.G., Schuyler, Q.A., Smith, S.D.A., Takada, H., Townsend, K.A., Wabnitz, C.C.C., Young, L.C. & Hamann, M. 2014. Hamman, M. 2014. Global research priorities to mitigate plastic pollution impacts on marine wildlife. Endangered Species Research 25: 225-247.

Ryan, P.G. 2013. A simple technique for counting marine debris at sea reveals steep litter gradients between the Straits of Malacca and the Bay of Bengal. Marine Pollution Bulletin 69: 128-136.

Ryan, P.G., Bouwman, H., Moloney, C.L., Yuyama, M. & Takada, H. 2012. Long-term decreases in persistent organic pollutants in South African coastal waters detected from beached polyethylene pellets. Marine Pollution Bulletin 64: 2756-2760.

Ryan, P.G., Moore, C.J., van Franeker, J.A. & Moloney, C.L. 2009. Monitoring the abundance of plastic debris in the marine environment. Philosophical Transactions of the Royal Society B 364: 1999-2012.

Ryan, P.G. 2008. Seabirds indicate decreases in plastic pellet litter in the Atlantic and south-western Indian Ocean. Marine Pollution Bulletin 56: 1406-1409.

Cliff, G., Dudley, S.F.J., Ryan, P.G. & Singleton, N. 2002. Large sharks and plastic debris in KwaZulu-Natal, South Africa. Marine and Freshwater Research 53: 575-581.

Ballance, A., Ryan, P.G. & Turpie, J.K. 2000. How much is a clean beach worth? The impact of litter on beach users in the Cape Peninsula, South Africa. South African Journal of Science 96: 210-213.

Ryan, P.G. 1996. Plastic litter in marine systems: impacts, sources and solutions. Plastics Southern Africa 26 (6): 20-28.

Ryan, P.G. 1996. Policy recommendations for controlling plastic litter. Plastics Southern Africa 25 (10): 46.

Ryan, P.G. & Swanepoel, D. 1996. Cleaning beaches: sweeping litter under the carpet. South African Journal of Science 92: 275-276.

Ryan, P.G., Swanepoel, D., Rice, N. & Preston, G. 1996. The ‘free’ shopping bag debate: costs and attitudes. South African Journal of Science 92: 163-165.

Ryan, P.G. & Moloney, C.L. 1993. Marine litter keeps increasing. Nature 361: 23.

Ryan, P.G. & Moloney, C.L. 1990. Plastic and other artefacts on South African beaches: temporal trends in abundance and composition. South African Journal of Science 86: 450-452.

Ryan, P.G. 1988. Intraspecific variation in plastic ingestion by seabirds and the flux of plastic through seabird populations. Condor 90: 446-452.

Ryan, P.G. 1988. The characteristics and distribution of plastic particles at the sea-surface off the southwestern Cape Province, South Africa. Marine Environmental Research 25: 249-273.

Ryan, P.G. 1988. Effects of plastic ingestion on seabird feeding: evidence from chickens. Marine Pollution Bulletin 19: 125-128.

Ryan, P.G., Connell, A.D. & Gardner, B.D. 1988. Plastic ingestion and PCBs in seabirds: is there a relationship? Marine Pollution Bulletin 19: 174-176.

Ryan, P.G. & Fraser, M.W. 1988. The use of Great Skua pellets as indicators of plastic pollution in seabirds. Emu 88: 16-19.

Ryan, P.G. & Watkins, B.P. 1988. Accumulation of stranded plastic objects and other artefacts at Inaccessible Island, central South Atlantic Ocean. South African Journal of Antarctic Research 18: 11-13.

Ryan, P.G. 1987. The incidence and characteristics of plastic particles ingested by seabirds. Marine Environmental Research 23: 175-206.

Ryan, P.G. 1987. The origin and fate of artefacts stranded on islands in the African sector of the Southern Ocean. Environmental Conservation 14: 341-346.

Ryan, P.G. 1987. The effects of ingested plastic on seabirds: correlations between plastic load and body condition. Environmental Pollution 46: 119-125.

Ryan, P.G. & Jackson, S. 1987. The lifespan of ingested plastic particles in seabirds and their effect on digestive efficiency. Marine Pollution Bulletin 18: 217-219.

Ryan, P.G. 1986. Seabirds as water quality monitors. Marine Pollution Bulletin 17: 435-436.

Seabird-fishery interactions and other bycatch issues (38 papers)

Rollinson, D.P., Dilley, B.J. & Ryan, P.G. 2014. Diving behaviour of white-chinned petrels and its relevance for mitigating longline bycatch. Polar Biology 37: 1301-1308.

Glass, J.P. & Ryan, P.G. 2013. Reduced seabird night strikes and mortality in the Tristan rock lobster fishery. African Journal of Marine Science 35: 589-592.

Reid, T.A., Wanless, R.M., Hilton, G.M., Phillips, R.A. & Ryan, P.G. 2013. Foraging range and habitat associations of non-breeding Tristan Albatrosses: overlap with fisheries and implications for conservation. Endangered Species Research 22: 39-49.

Tew Kai, E., Benhamou, S., van der Lingen, C., Coetzee, J., Pichegru, L., Ryan, P.G. & Grémillet, D. 2013. Are Cape gannets dependent upon fishery waste in the southern Benguela upwelling system? A multi-scale analysis using seabird GPS-tracking, hydro-acoustic estimates of pelagic fish, and vessel monitoring systems. Journal of Applied Ecology 50: 659-670.

Pichegru, L., Ryan, P.G., van Eeden, R., Reid, T., Grémillet, D. & Wanless, R. 2012. Industrial fishing, no-take zones and endangered penguins. Biological Conservation 156: 117-125.

Grémillet, D., Mullers, R.H.E., Moseley, C., Pichegru, L., Coetzee, J.C., Sabarros, P.S., van der Lingen, C., Ryan, P.G., Kato, A. & Ropert-Coudert, Y. 2010. Seabirds, fisheries and cameras. Frontiers in Ecology and the Environment 8: 401-402.

Pichegru, L., Grémillet, D., Crawford, R.J.M., & Ryan, P.G. 2010. Marine no-take zone rapidly benefits threatened penguin. Biology Letters 6: 498-501.

Petersen, S.L., Honig, M.B., Nel, R., Ryan, P.G. & Underhill, L.G. 2009. Turtle bycatch in the pelagic longline fishery off southern Africa. African Journal of Marine Science 31: 87-96.

Petersen, S.L., Honig, M.B., Ryan, P.G. & Underhill, L.G. 2009. Seabird bycatch in the pelagic longline fishery off southern Africa. African Journal of Marine Science 31: 191-204.

Petersen, S.L., Honig, M.B., Ryan, P.G., Underhill, L.G. & Goren, M. 2009. Seabird bycatch in the demersal longline fishery off southern Africa. African Journal of Marine Science 31: 205-214.

Petersen, S.L., Honig, M.B., Ryan, P.G., Underhill, L.G. & Compagno, L.J.V. 2009. Pelagic shark bycatch in the tuna- and swordfish-directed longline fishery off southern Africa. African Journal of Marine Science 31: 215-225.

Pichegru, L., Ryan, P.G., Le Bohec, C., van der Lingen, C.D., Navarro, R., Petersen, S., Lewis, S., van der Westhuizen, J. & Grémillet, D. 2009. Overlap between vulnerable top predators and fisheries in the Benguela upwelling system: implications for marine protected areas. Marine Ecology Progress Series 391: 199-208.

Grémillet, D., Pichegru, L., Kuntz, G., Woakes, A.G., Wilkinson, S., Crawford, R.J.M. & Ryan, P.G. 2008. A junk-food hypothesis for gannets feeding on fishery waste. Proceedings of the Royal Society London B 275: 1149-1156.

Abbott, C.L., Double, M.C., Gales, R., Baker, G.B., Lashko, A., Robertson, C.J.R. & Ryan, P.G. 2006. Molecular provenance analysis for Shy and White-capped Albatrosses killed by fisheries interactions in Australia, New Zealand and South Africa. Conservation Genetics 7: 531-542.

Ryan, P.G. & Watkins, B.P. 2008. Estimating seabird bycatch by fisheries and its impact on seabird populations. Animal Conservation 11: 260-262.

Ryan, P.G. & Watkins, B.P. 2008. Estimating seabird bycatch by fisheries and its impact on seabird populations. Animal Conservation 11: 260-262.

Petersen, S.L., Phillips, R., Ryan, P.G. & Underhill, L.G. 2008. Albatross overlap with fisheries in the Benguela Upwelling System: implications for conservation and management. Endangered Species Research 5: 117-127.

Watkins, B.P., Petersen, S.L. & Ryan, P.G. 2008. Interactions between seabirds and deep-water hake trawl gear: an assessment of impacts in South African waters. Animal Conservation 11: 247-254.

Baker, G.B., Double, M.C., Gales, R., Tuck, G.N., Abbott, C.L., Ryan, P.G., Petersen, S.L, Robertson, C.J.R. & Alderman, R. 2007. A global assessment of the impact of fisheries-related mortality on Shy and White-capped albatrosses: conservation implications. Biological Conservation 137: 319-333.

Ryan, P.G., Nel, D.C. & Cooper, J. 2006. Reducing longline mortality of seabirds: a South African perspective. Acta Zoologica Sinica 52S: 645-648.

Cuthbert, R.J., Hilton, G., Ryan, P.G. & Tuck, G. 2005. At-sea distribution of the Tristan Albatross Diomedea dabbenena and potential interactions with pelagic longline fishing in the South Atlantic Ocean. Biological Conservation 121: 345-355.

Mills, M.S.L. & Ryan, P.G. 2005. Modelling impacts of long-line fishing: what are the effects of pair-bond disruption and sex-biased mortality on albatross fecundity? Animal Conservation 8: 359-367.

Nel, D.C., Taylor, F., Ryan, P.G. & Cooper, J. 2003. Population dynamics of Wandering Albatrosses Diomedea exulans at sub-Antarctic Marion Island: long-line fishing and environmental influences. African Journal of Marine Science 25: 503-517.

Ryan, P.G., Keith, D.G. & Kroese, M. 2002. Seabird bycatch by tuna longline fisheries off southern Africa. South African Journal of Marine Science 24: 103-110.

Ryan, P.G. & Watkins, B.P. 2002. Reducing incidental mortality of seabirds with an underwater longline setting funnel. Biological Conservation 104: 127-131.

Nel, D.C., Ryan, P.G., Nel, J.L., Klages, N.T.W., Wilson, R.P., Robertson, G. & Tuck, G.N. 2002. Foraging interactions between Wandering Albatrosses Diomedea exulans breeding on Marion Island and long-line fisheries in the southern Indian Ocean. Ibis 144: E141-E154.

Nel, D.C., Ryan, P.G. & Watkins, B.P. 2002. Seabird mortality in the Patagonian toothfish longline fishery around the Prince Edward Islands, 1996-2000. Antarctic Science 14: 151-161.

Nel, D.C., Ryan, P.G., Crawford, R.J.M., Cooper, J. & Huyser, O.A.W. 2002. Population trends of albatrosses and petrels breeding at sub-Antarctic Marion Island. Polar Biology 25: 81-89.

Glass, N., Lavarello, I., Glass, J.P. & Ryan, P.G. 2000. Longline fishing at Tristan da Cunha: impacts on seabirds. Atlantic Seabirds 2: 49-56.

Nel, D.C., Nel, J.L., Ryan, P.G., Klages, N.T.W., Wilson, R.P. & Robertson, G. 2000. The foraging ecology of Grey-headed Mollymawks at Marion Island, southern Indian Ocean, in relation to known longline fishing activity. Biological Conservation 96: 219-231.

Ryan, P.G. 1999. Sexual dimorphism, moult and body condition of seabirds killed by longline vessels around the Prince Edward Islands, 1996-97. Ostrich 70: 187-192.

Ryan, P.G. & Boix-Hinzen, C. 1999. Consistent male-biased seabird mortality in the Patagonian Toothfish long-line fishery. Auk 116: 851-854.

Ryan, P.G. & Boix-Hinzen, C. 1998. Tuna longline fisheries off southern Africa: the need to limit seabird bycatch. South African Journal of Science 94: 179-182.

Barnes, K.N., Ryan, P.G. & Boix-Hinzen, C. 1997. The impact of the hake Merluccius spp. longline fishery off South Africa on procellariiform seabirds. Biological Conservation 82: 227-234.

Moloney, C.L., Cooper, J., Ryan, P.G. & Siegfried, W.R. 1994. Use of a population model to assess the impact of longline fishing on Wandering Albatross Diomedea exulans populations. Biological Conservation 70: 195-204.

Ryan, P.G. 1991. The impact of the commercial lobster fishery on seabirds at the Tristan da Cunha islands, South Atlantic Ocean. Biological Conservation 57: 339-350.

Ryan, P.G. & Cooper, J. 1991. Rockhopper Penguins and other marine life threatened by driftnet fisheries at Tristan da Cunha. Oryx 25: 76-79.

Ryan, P.G. & Moloney, C.L. 1988. The effect of trawling on bird and seal distributions in the southern Benguela region. Marine Ecology Progress Series 45: 1-11.

Seabird monitoring and conservation (34 papers)

Pichegru, L., Edwards, T.B., Dilley, B.J., Flower, T.P. & Ryan, P.G. 2015. African Penguin tolerance to humans depends on historical exposure at colony level. Bird Conservation International

Whittington, P.A., Crawford, R.J.M., Martin, A.P., Randall, R.M., Brown, M., Makhado, A.B., Dyer, B.M., Upfold, L., Harrison, K.H., Huisamen, J., Ryan, P.G., Waller, L. & Witteveen, M. in press. Recent trends of the Kelp Gull Larus dominicanus in South Africa. Waterbirds

Cuthbert, R.J., Cooper, J. & Ryan, P.G. 2014. Population trends and breeding success of albatrosses and giant petrels at Gough Island in the face of at-sea and on-land threats. Antarctic Science 26: 163-171.

Rexer-Huber, K., Parker, G.C., Ryan, P.G. & Cuthbert, R.J. 2014. Burrow occupancy and population size in the Atlantic Petrel: a comparison of methods. Marine Ornithology 42: 137-141.

Crawford, R.J.M., Randall, R.M., Whittington, P.A., Waller, L., Dyer, B.M., Allan, D.G., Fox, C., Martin, A.P., Upfold, L., Visagie, J., Bachoo, S., Bowker, M., Downs, C.T., Fox, R., Huisamen, J., Makhado, A.B., Oosthuizen, W.H., Ryan, P.G., Taylor, R.H. & Turpie, J.K. 2013. South Africa's coastal-breeding White-breasted Cormorants: population trends, breeding season and movements, and diet. African Journal of Marine Science 35: 473-490.

Ryan, P.G. & Kerr, J. 2012. Is fur seal predation driving the decrease in Northern Rockhopper Penguins Eudyptes moseleyi at Gough Island? Marine Ornithology 40: 73-75.

Ryan, P.G., Dilley, B.J. & Jones, M.G.W. 2012. The distribution and abundance of White-chinned Petrels (Procellaria aequinoctialis) breeding at the sub-Antarctic Prince Edward Islands. Polar Biology 35: 1851-1859.

Lewis, S.F.F., Turpie, J.K. & Ryan, P.G. 2012. Are African Penguins worth saving? The ecotourism value of the Boulders Beach colony. African Journal of Marine Science 34: 497-504.

Lewison, R., Oro, D., Godley, B., Underhill, L., Bearhop, S., Wilson, R., Ainley, D., Arcos, J.M., Boersma, P.D., Borboroglu, P., Boulinier, T., Frederiksen, M., Genovart, M., González-Solís, J., Green, J.A., Grémillet, D., Hamer, K.C., Hilton, G.M., Hyrenbach, K.D., Martínez-Abraín, A., Montevecchi, W.A., Phillips, R.A., Ryan, P.G., Sagar, P., Sydeman, W.J., Yorio, P., Wanless, S., Watanuki, Y. & Weimerskirch, H. 2012. Research priorities for seabirds: improving seabird conservation and management in the 21st century. Endangered Species Research 17: 93-121.

Ryan, P.G. & Ronconi, R.A. 2011. Continued increase in numbers of Spectacled Petrels Procellaria conspicillata. Antarctic Science 23: 332-336.

Crawford, R.J.M., Altwegg, R., Barham, B.J., Barham, P.J., Durant, J.M., Dyer, B.M., Geldenhuys, D., Makhado, A.B., Pichegru, L., Ryan, P.G., Underhill, L.G., Upfold, L., Visagie, J., Waller, L.J. & Whittington, P.A. 2011. Collapse of South Africa’s penguins in the early 21st century. African Journal of Marine Science 33: 139-156.

Robson, B., Glass, T., Glass, N., Glass, J., Green, J., Repetto, C., Rodgers, G., Ronconi, R.A., Ryan, P.G., Swain, G. & Cuthbert, R.J. 2011. Revised population estimate and trends for the endangered Northern Rockhopper Penguin Eudyptes moseleyi at Tristan da Cunha. Bird Conservation International 21: 454-459.

Mwema, M.M., de Ponte Machado, M. & Ryan, P.G. 2010. Breeding seabirds at Dassen Island, South Africa: chances of surviving Great White Pelican predation. Endangered Species Research 9: 125-131.

Pichegru, L., Ryan, P.G., Crawford, R.J.M., van der Lingen, C. & Grémillet, D. 2010.  Behavioural inertia places a top marine predator at risk from environmental change in the Benguela upwelling system. Marine Biology 157: 537-544.

Ryan, P.G., Jones, M.G.W., Dyer, B.M., Upfold, L. & Crawford, R.J.M. 2009. Recent population estimates and trends in numbers of albatrosses and giant petrels breeding at the sub-Antarctic Prince Edward Islands. African Journal of Marine Science 31: 409-417.

Ryan, P.G., Whittington, P.A. & Crawford, R.J.M. 2009. A tale of two islands: contrasting fortunes for Subantarctic Skuas at the Prince Edward Islands. African Journal of Marine Science 31: 431-437.

Converse, S.J., Kendall, W.L., Doherty, P.F.Jr. & Ryan, P.G. 2009. Demography of Marion Island Grey-headed Albatross: analysis utilizing advances in mark-recapture models. Auk 126: 77-88.

Crawford, R.J.M., Ryan, P.G., Dyer, B.M. & Upfold, L. 2009. Recent trends in numbers of Crozet Shags breeding at the Prince Edward Islands. African Journal of Marine Science 31: 427-430.

Crawford, R.J.M., Whittington, P.A., Upfold, L., Ryan, P.G., Petersen, S.L., Dyer, B.M. & Cooper, J. 2009. Recent trends in numbers of four species of penguins at the Prince Edward Islands. African Journal of Marine Science 31: 419-426.

Cuthbert, R., Cooper, J., Burle, M.-H., Glass, C.J., Glass, J.P., Glass, T., Hilton, G.M., Sommer, E., Wanless, R.M. & Ryan, P.G. 2009. Population trends and conservation status of the Northern Rockhopper Penguin Eudyptes moseleyi at Tristan da Cunha and Gough Island. Bird Conservation International 19: 109-120.

Hampton, S., Ryan, P.G. & Underhill, L.G. 2009. The effect of flipper banding on the breeding success of African Penguins Spheniscus demersus at Boulders Beach, South Africa. Ostrich 80: 77-80.

Whittington, P.A., Crawford, R.J.M., Dyer, B.M. & Ryan, P.G. 2009. Estimates of numbers of kelp gulls and Kerguelen and Antarctic terns breeding at the Prince Edward Islands, 1996/97–2008/09. African Journal of Marine Science 31: 439-444.

Ryan, P.G., Dorse, C. & Hilton, G.M. 2006. The conservation status of the Spectacled Petrel Procellaria conspicillata. Biological Conservation 131: 575-583.

Lewis, S., Daunt, F., Grémillet, D., Ryan, P.G., Crawford, R.J.M. & Wanless, S. 2006. Using behavioural variables to identify proximate causes of population change in a seabird. Oecologia 147: 606-614.

Cuthbert, R.J., Sommer, E.S., Ryan, P.G., Cooper, J., & Hilton, G. 2004. Demography and conservation status of the Tristan Albatross Diomedea [exulans] dabbenena. Biological Conservation 117: 471-481.

Ryan, P.G., Cooper, J., Dyer, B.M., Underhill, L.G., Crawford, R.J.M. & Bester, M.N. 2003. Counts of surface-nesting seabirds breeding at sub-Antarctic Prince Edward Island, summer 2001/02. African Journal of Marine Science 25:  441-451.

Crawford, R.J.M., Cooper, J., Dyer, B.M., Greyling, M.D., Klages, N.T.W., Ryan, P.G., Petersen, S.L., Underhill, L.G., Upfold, L., Wilkinson, W., de Villiers, M.S., du Plessis, S., du Toit, M., Leshoro, T.M., Makhado, A.B., Mason, M., Merkle, D., Tshingana, D., Ward, V.L. & Whittington, P.A. 2003. Populations of surface-nesting seabirds at Marion Island, 1994/95-2002/03. African Journal of Marine Science 25: 427-439.

Cuthbert, R.J., Ryan, P.G., Cooper, J. & Hilton, G. 2003. Demography and population trends of the Atlantic Yellow-nosed Albatross Thalassarche chlororhynchos. Condor 105: 439-452.

Ryan, P.G., Avery, G. & Steele, W.K. 1991. Distribution, population sizes and conservation of marine and coastal birds in False Bay. Transactions of the Royal Society of South Africa 47: 649-662.

Ryan, P.G., Dean, W.R.J., Moloney, C.L., Watkins, B.P. & Milton, S.J. 1990. New information on seabirds at Inaccessible Island and other islands in the Tristan da Cunha group. Marine Ornithology 18: 43-54.

Ryan, P.G. & Watkins, B.P. 1988. Birds of the inland mountains of western Dronning Maud Land, Antarctica. Cormorant 16: 34-40.

Brooke, R.K., Cooper, J., Hockey, P.A.R., Ryan, P.G., Sinclair, J.C., Suter, W. & Tree, A.J. 1988. Distribution, population size and conservation of the Antarctic Tern Sterna vittata in southern Africa. Cormorant 16: 107-113.

Fraser, M.W., Ryan, P.G. & Watkins, B.P. 1988. The seabirds of Inaccessible Island, South Atlantic Ocean. Cormorant 16: 7-33.

Ryan, P.G. 1987. The distribution, population size and foraging behaviour of Kerguelen Terns at the Prince Edward Islands. South African Journal of Antarctic Research 17: 163-166.

Foraging ecology of seabirds and other marine predators (42 papers)

Massie, P., McIntyre, T., Ryan, P.G., Bester, M.N., Bornemann, H., Ansorge, I. in press. The role of eddies on the diving behaviour of female Southern Elephant Seals. Polar Biology

McInnes, A.M., Khoosal, A., Murrell, B., Merkle, D., Lacerda, M., Nyengera, R., Coetzee, J.C., Edwards, L.C., Ryan, P.G., Rademan, J., van der Westhuizen, J.J. & Pichegru, L. in press. Recreational fish-finders - an inexpensive alternative to scientific echo-sounders for unravelling the links between marine top predators and their prey. PLoS ONE 10/2015

van Eeden, R., Reid, T.A., Ryan, P.G. & Pichegru, L. in press. Fine-scale foraging cues for African Penguins in a highly variable marine environment. Marine Ecology Progress Series

Whitehead, T.O., Kato, A., Ropert-Coudert, Y. and Ryan, P.G. in press. Habitat use and diving behaviour of Macaroni Eudyptes chrysolophus and Eastern Rockhopper E. chrysocome filholi penguins during the critical pre-moult period. Marine Biology

Cohen, L.A., Pichegru, L., Grémillet, D., Coetzee, J., Upfold, L. & Ryan, P.G. 2014. A decade of changes in prey availability impact the foraging behaviour and fitness of Cape Gannets. Marine Ecology Progress Series 505: 281-293.

Reid, T.A., Ronconi, R.A., Cuthbert, R.J. & Ryan, P.G. 2014. The summer foraging ranges of adult Spectacled Petrels Procellaria conspicillata. Antarctic Science 26: 23-32.

Dilley, B.J., Davies, D., Connan, M., Cooper, J., de Villiers, M., Swart, L., Vandenabeele, S., Ropert-Coudert, Y. & Ryan, P.G. 2013. Giant petrels as predators of albatross chicks. Polar Biology 36: 761-766.

Ryan, P.G., Edwards, L. & Pichegru, L. 2012. African Penguins Spheniscus demersus, bait balls and the Allee effect. Ardea 100: 89-94.

Cook, T.R., Hamann, M., Pichegru, L., Bonadonna, F., Grémillet, D. & Ryan, P.G. 2012. GPS and time-depth loggers reveal underwater foraging plasticity in a flying diver, the Cape Cormorant. Marine Biology 159: 379-387.

Moseley, C., Grémillet, D., Connan, M., Ryan, P.G., Mullers, R.H.E., van der Lingen, C.D., Coetzee, J.C., Crawford, R.J.M., Sabarros, P., McQuaid, C. & Pichegru, L. 2012. Foraging ecology and ecophysiology of Cape gannets from colonies with contrasting environments and population dynamics. Journal of Experimental Marine Biology and Ecology 422-423: 29-38.

Pichegru, L., Ropert-Coudert, Y., Kato, A., Takahashi, A., Dyer, B.M. & Ryan, P.G. 2011. Diving patterns of female Macaroni Penguins breeding on Marion Island, South Africa. Polar Biology 34: 945-954.

Wright, K.L.B., Pichegru, L. & Ryan, P.G. 2011. Penguins are attracted to dimethyl sulphide at sea. Journal of Experimental Biology 214: 2509-2511.

Ryan, P.G., Pichegru, L., Ropert-Coudert, Y., Grémillet, D. & Kato, A. 2010. On a wing and a prayer: the foraging ecology of breeding Cape Cormorants. Journal of Zoology London 280: 25-32.

Ronconi, R.A., Ryan, P.G. & Ropert-Coudert, Y. 2010. Diving of Great Shearwaters (Puffinus gravis) in cold and warm water regions of the South Atlantic Ocean. PLoS ONE 5(11): e15508. doi:10.1371/journal.pone.0015508.

Wilson, J.W., Burle, M-H., Cuthbert, R., Stirnemann, R.L. & Ryan, P.G. 2010. Breeding success of Northern Rockhopper Penguins (Eudyptes moseleyi) at Gough Island, South Atlantic Ocean. Emu 110: 137-141.

Ryan, P.G. 2009. The seabirds of Conrad Rise, Southern Ocean. Bulletin of the African Bird Club 16: 67-77.

Ropert-Coudert, Y., Daunt, F., Kato, A., Ryan, P.G., Lewis, S., Kobayashi, K., Mori, Y., Grémillet, D. & Wanless, S. 2009. Underwater wingbeats extend depth and duration of plunge dives in gannets. Journal of Avian Biology 40: 380-387.

Ryan, P.G., Sommer, E. & Breytenbach, E. 2008. Giant petrels Macronectes hunting Northern Rockhopper Penguins Eudyptes moseleyi at sea. Ardea 96: 129-134.

Cunningham, G.B., Strauss, V. & Ryan, P.G. 2008. African Penguins (Spheniscus demersus) can detect dimethyl sulphide, a prey-related odour. Journal of Experimental Biology 211: 32123-3127.

Grémillet, D., Lewis, S., Drapeau, L., van der Lingen, C.D., Huggett, J.A., Coetzee, J.C., Verheye, H.M., Daunt, F., Wanless, S. & Ryan, P.G. 2008. Spatial match-mismatch in the Benguela upwelling zone: should we expect chlorophyll and SST to predict marine predator distributions? Journal of Applied Ecology 45: 610-621.

Ryan, P.G. 2007. Diving in shallow water: the foraging ecology of darters (Anhingidae). Journal of Avian Biology 38: 507-514.

Ryan, P.G., Petersen, S.L., Simeone, A. & Grémillet, D. 2007. Diving behaviour of African penguins: do they differ from other Spheniscus penguins? African Journal of Marine Science 29: 153-160.

Pichegru, L., Ryan, P.G., van der Lingen, C.D., Coetzee, J., Ropert-Coudert, Y. & Grémillet, D. 2007. Foraging behaviour and energetics of Cape Gannets Morus capensis feeding on live prey and fishery waste in the Benguela upwelling system. Marine Ecology Progress Series 350: 127-136.

Petersen, S.L., Ryan, P.G. & Grémillet, D. 2006. Is food availability limiting African Penguins at Boulders Beach? A comparison of foraging ecology at mainland and island colonies. Ibis 148: 14-26.

Ryan, P.G., Petersen, S.L., Peters, G. & Grémillet, D. 2004. GPS tracking a marine predator: the effects of precision, resolution and sampling rate on foraging tracks of African Penguins. Marine Biology 145: 215-223.

Grémillet, D., Dell’Omo, G., Ryan, P.G., Peters, G., Ropert-Coudert, Y. & Weeks, S.J. 2004. Offshore diplomacy, or how seabirds mitigate intra-specific competition: A case study based on GPS tracking of Cape gannets from neighbouring breeding sites. Marine Ecology Progress Series 268: 265-279.

Ropert-Coudert, Y., Grémillet, D., Kato, A., Ryan, P.G., Naito, Y. & Le Maho, Y. 2004. A fine-scale time budget of Cape Gannets provides insights into the foraging strategies of coastal seabirds. Animal Behaviour 67: 985-992.

Ropert-Coudert, Y., Grémillet, D., Ryan, P.G., Kato, A., Naito, Y. & Le Maho, Y. 2004. Between air and water: the gannet’s plunge dive. Ibis 146: 281-290.

Nel, D.C., Lutjeharms J.R.E., Pakhomov, E.A., Ansorge, I.J., Ryan, P.G. & Klages, N.T.W. 2001. Exploitation of mesoscale oceanographic features by Grey-headed Albatrosses Thalassarche chrysostoma in the southern Indian Ocean. Marine Ecology Progress Series 217: 15-26.

Ryan, P.G. & Nel, D.C. 1999. Foraging behaviour of diving petrels. Emu 99: 72-74.

Wilson, R.P., Hustler, K., Ryan, P.G., Burger, A.E. & Nöldeke, E.C. 1992. Diving birds in cold water: do Archimedes and Boyle determine energetic costs? American Naturalist 140: 179-200.

Ryan, P.G. & Moloney, C.L. 1991. Prey selection and temporal variation in the diet of Subantarctic Skuas at Inaccessible Island, Tristan da Cunha. Ostrich 62: 52-58.

Crawford, R.J.M., Ryan, P.G. & Williams, A.J. 1991. Seabird consumption and production in the Benguela and western Agulhas ecosystems. South African Journal of Marine Science 11: 357-375.

Hockey, P.A.R., Ryan, P.G. & Bosman, A.L. 1989. Age-related intraspecific kleptoparasitism and foraging success of Kelp Gulls Larus dominicanus. Ardea 77: 205-210.

Wilson, R.P., Ryan, P.G. & Wilson, M.-P. 1989. Sharing food in the stomachs of seabirds between adults and chicks - a case for delayed gastric emptying. Comparative Biochemistry & Physiology 94A: 461-466.

Hockey, P.A.R., Bosman, A.L. & Ryan, P.G. 1988. What determines prey selection by Kelp Gulls Larus dominicanus in multispecies mussel communities? Cormorant 16: 103-106.

Ryan, P.G. 1987. The foraging behaviour and breeding seasonality of Hartlaub's Gull Larus hartlaubii. Cormorant 15: 23-32.

Ryan, P.G., Hockey, P.A.R. & Bosman, A.L. 1987. The foraging behavior of Gray Gulls at a sandy beach. Wilson Bulletin 99: 271-273.

Ryan, P.G., Wilson, R.P. & Cooper, J. 1987. Intraspecific mimicry and status signals in juvenile African Penguins. Behavioural Ecology & Sociobiology 20: 69-76.

Wilson, R.P., Ryan, P.G., James, A. & Wilson, M.-P. 1987. Conspicuous coloration may enhance prey capture in some piscivores. Animal Behaviour 35: 1558-1560.

Ryan, P.G. & Jackson, S. 1986. Stomach pumping: is killing seabirds necessary? Auk 103: 427-428.

Jackson, S. & Ryan, P.G. 1986. Differential digestion rates of prey by White-chinned Petrels. Auk 103: 617-619.

Sundry seabird papers (35 papers)

Ryan, P.G., Ferreira, C., Perold, V., Osborne, A. & Jones, C.W. in press. Failure to launch: evidence of protracted parental care in albatrosses. Seabird

Ryan, P.G., Dilley, B.J., Jones, C. & Bond, A. 2015. Blue Petrels breeding on Gough Island. Ostrich 89: 193-194. (doi: 10.2989/00306525.2015.1005558)

Grecian, J., McGill, R., Phillips, R.A., Ryan, P.G. & Furness, R.W. 2015. Quantifying variation in δ13C and δ15N isotopes within and between feathers and individuals; is one sample enough? Marine Biology 162: 733-741. doi: 10.1007/s00227-015-2618-8

Jones, C.W., Risi, M.M., Kuntz, W., Ryan, P.G., Steinfurth, A. & Bond, A.L. 2015. Bill deformities in penguins (Spheniscidae): a global review. Marine Ornithology 43: 207–209.

Torres, L.G., Sutton, P.J., Thomspon, D.R., Delord, K., Weimerskirch, H., Sagar, P.M., Sommer, E., Dilley, B.J., Ryan, P.G. & Phillips, R.A. 2015. Poor transferability of species distribution models for a pelagic predator, the Grey Petrel, indicates contrasting habitat preferences across ocean basins. PLoS ONE 10(3): e0120014.

Witteveen, M., Brown, M. & Ryan, P.G. 2015. Pseudo-egg and exotic egg adoption by Kelp Gulls Larus dominicanus vetula. African Zoology 50: 59-61.

Colabuono, F.I., Barquete, V., Taniguchi, S., Ryan, P.G. & Montone, R.C. 2014. Stable isotopes of carbon and nitrogen in the study of organochlorine contaminants in albatrosses and petrels. Marine Pollution Bulletin 83: 241-247.

Commins, M.L., Ansorge, I. & Ryan, P.G. 2014. Multi-scale factors influencing seabird assemblages in the African sector of the Southern Ocean. Antarctic Science 26: 38-48.

Jones, M.G.W. & Ryan, P.G. 2014. Effects of pre-laying attendance and body condition on long-term reproductive success in Wandering Albatrosses. Emu 114: 137-145.

Jones, M.G.W., Dilley, B.J., Hagens, Q.A., Louw, H., Mertz, E.M., Visser, P. & Ryan, P.G. 2014. The effect of parental age, experience and historic reproductive success on Wandering Albatross (Diomedea exulans) chick growth and survival. Polar Biology 37: 1633-1644.

Ryan, P.G. 2013. Moult of flight feathers in darters (Anhingidae). Ardea 101: 177-180.

Ryan, P.G., Dyer, B.M., Martin, A.P., Ward, V.L. & Whittington, P.A. 2013. Supernormal clutches in southern African Kelp Gulls Larus dominicanus vetula. Ostrich 84: 157-160.

Ryan, P.G., Rose, B., Carter, M. & Clarke, R.H. 2013. A review of Jouanin’s Petrel records in the eastern Indian Ocean. Ostrich 84: 157-160.

Abad, F.X., Busquets, N., Sánchez, A., Ryan, P.G., Majó, N. & González-Solís, J. 2013. Serological and virological surveys of Influenza A virus in Antarctic and sub-Antarctic penguins. Antarctic Science 25: 339-344.

Barquete, V., Strauss, V. & Ryan, P.G. 2013. Stable isotope turnover in blood and claws: a case study in captive African Penguins. Journal of Experimental Marine Biology and Ecology 448: 121-127.

Jones, M.G.W., Techow, N.M.S.M. & Ryan, P.G. 2012. Dalliances and doubtful dads: What determines extra-pair paternity in socially monogamous Wandering Albatrosses? Behavioural Ecology & Sociobiology 66: 1213-1224.

Ryan, P.G., Cuthbert, R.J. & Cooper, J. 2007. Two egg clutches among albatrosses. Emu 107: 210-213.

Ryan, P.G., Phillips, R.A., Nel, D.C. & Wood, A.G. 2007. Breeding frequency in Grey-headed Albatrosses. Ibis 149: 45-52.

Ropert-Coudert, Y., Wilson, R.P., Grémillet, D., Kato, A., Lewis, S. & Ryan, P.G. 2006. Electrocardiogram recordings in free-ranging gannets reveal minimum difference in heart rate during flapping versus gliding flight. Marine Ecology Progress Series 328: 275-284.

de Villiers, M.S., Cooper, J. & Ryan, P.G. 2005. Individual variability of behavioural responses by Wandering Albatrosses (Diomedea exulans) to human disturbance. Polar Biology 28: 255-260.

Ropert-Coudert, Y., Grémillet, D., Gachot-Neveu, H., Lewis, S. & Ryan, P.G. 2005. Seeking dimorphism in monomorphic species: the lure of the gannet’s mask. Ostrich 76: 212-214.

Ryan, P.G. 2004. Pink-backed Pelicans feeding communally with Great White Pelicans. Ostrich 75: 168.

Cuthbert, R.J., Phillips, R.A. & Ryan, P.G. 2003. Separating the Tristan Albatross and the Wandering Albatross using morphometric measurements. Waterbirds 26: 338-344.

Keith, D.G., Harck, B.I.B., Ryan, P.G. & Mehlum, F. 2002. Post-breeding dispersal of Northern giant Petrels Macronectes halli from Marion to Bouvet Islands. Marine Ornithology 30: 31.

Ryan, P.G. 2001. Partial leucism in Whitechinned Petrels. Bird Numbers 10(2): 6-7.

Ryan, P.G., Steele, W.K., Siegfried, W.R. & Vogel, J.C. 1992. Radiocarbon dates of Snow Petrel regurgitations can reveal exposure periods for nunataks in Antarctica. South African Journal of Science 88: 578-580.

Ryan, P.G, Avery, G., Rose, B., Ross, G.J.B., Sinclair, J.C. & Vernon, C.J. 1989. The Southern Ocean seabird irruption to South African waters during winter 1984. Cormorant 17: 41-55.

Ryan, P.G. & Cooper, J. 1989. The distribution and abundance of aerial seabirds in relation to Antarctic krill in the Prydz Bay region, Antarctica, during late summer. Polar Biology 10: 199-209.

Ryan, P.G. & Cooper, J. 1989. Observer precision and bird conspicuousness during counts of birds at sea. South African Journal of Marine Science 8: 271-276.

Ryan, P.G. & Watkins, B.P. 1989. Snow Petrel breeding biology at an inland site in continental Antarctica. Colonial Waterbirds 12: 176-184.

Hoberg, E.P. & Ryan, P.G. 1989. Ecology of helminth parasitism in Puffinus gravis (Procellariiformes) on the breeding grounds at Gough Island. Canadian Journal of Zoology 67: 220-225.

Duffy, D.C., Ryan, P.G., Wilson, R.P. & Wilson, M.-P. 1988. Spring seabird distribution in the Straits of Magellan. Cormorant 16: 98-102.

Ryan, P.G. & Avery, G. 1987. Wreck of juvenile Blackbrowed Albatrosses on the west coast of South Africa during storm weather. Ostrich 58: 139-140.

Ryan, P.G. 1986. Trematode anklets on procellariiform seabirds from southern Africa and the adjacent Southern Ocean. Cormorant 13: 157-161.

Ryan, P.G. & Hunter, S. 1985. Early breeding of Imperial Cormorants at Prince Edward Island. Cormorant 13: 31-34.

Island biology and conservation (42 papers)

Cerfonteyn, M. & Ryan, P.G. in press. Have burrowing petrels recovered on Marion Island two decades after cats were eradicated? Evidence from Subantarctic Skua prey remains. Antarctic Science doi: 10.1017/S0954102015000474

Cuthbert, R.J., Wanless, R.M., Angel, A., Burlé, M.-H., Hilton, G.M., Louw, H., Visser, P., Wilson, J.W. & Ryan, P.G. in press. Drivers of predatory behaviour and extreme body size in House Mice Mus musculus on Gough Island. Journal of Mammalogy

Dilley, B.J., Davies, D., Bond, A.L. & Ryan, P.G. in press. Effects of mouse predation on burrowing petrel chicks at Gough Island. Antarctic Science

Gray, M.M., Parmenter, M., Hogan, C., Ford, I., Cuthbert, R.J., Ryan, P.G., Broman, K.W. & Payseur, B.A. in press. Genetics of rapid and extreme size evolution in island mice. Genetics doi: 10.1534/genetics.115.177790

Davies, D., Dilley, B.J., Bond, A.L., Cuthbert, R.J. & Ryan, P.G. 2015. Trends and tactics of mouse predation on Tristan Albatross Diomedea dabbenena chicks at Gough Island, South Atlantic Ocean. Avian Conservation and Ecology 10(1): 5.

Dilley, B.J., Schoombie, S., Schoombie, J. & Ryan, P.G. 2015. ‘Scalping’ of albatross fledglings by introduced mice spreads rapidly at Marion Island. Antarctic Science doi:10.1017/S0954102015000486

Ryan, P.G., Ortmann, H.E. & Herian, K. 2014. The impact of introduced scale insects on Phylica arborea and the implications for Nesospiza finches at Tristan da Cunha. Biological Conservation 176: 48-53.

Cuthbert, R.J., Broome, K., Bradley, J. & Ryan, P.G. 2014. Evaluating the effectiveness of aerial baiting operations for rodent eradications on cliffs on Gough Island, Tristan da Cunha. Conservation Evidence 11: 25-28.

Cuthbert, R.J., Louw, H., Lurling, J., Parker, G., Rexer-Huber, K., Sommer, E., Visser, P. & Ryan, P.G. 2013. Low burrow occupancy and breeding success of burrowing petrels at Gough Island: a consequence of mouse predation. Bird Conservation International 23: 113-124.

Gray, M.M., Wegmann, D., Haasl, R.J., White, M.A., Gabriel, S.I., Searle, J.B., Cuthbert, R.J., Ryan, P.G. & Payseur, B.A. 2014. Demographic history of a recent invasion of House Mice on the isolated island of Gough. Molecular Ecology 23: 1923-1939.

Cooper, J., Cuthbert, R.J. & Ryan, P.G. 2013. An overlooked biosecurity concern: back-loading at islands supporting introduced rodents. Aliens 33: 28-31.

Le Roux, P.C., Ramaswielaa, T., Kalwijb, J.M., Shaw, J.D., Ryan, P.G., Treasure, A.M., McClelland, G.T.W., McGeoch, M.A., Chown, S.L. 2013. Human activities, propagule pressure and alien plants in the sub-Antarctic: tests of generalities and evidence in support of management. Biological Conservation 161: 18-27.

Ryan, P.G., Glass, J.P., Glass, T., Barendse, J. & Cuthbert, R.C. 2012. Eradication of New Zealand flax Phormium tenax on Inaccessible and Nightingale Islands, Tristan da Cunha. Conservation Evidence 9: 58-62.

Chown, S.L., Lee, J.E., Hughes, K.A., Barnes, J., Barrett, P.J.,Bergstrom, D.M., Convey, P., Cowan, D.A., Crosbie, K., Dyer, G., Frenot, Y., Grant, S.M., Herr, D., Kennicutt, M.C., Lamers, M., Murray, A., Possingham, H.P., Reid, K., Riddle, M., Ryan, P.G., Sanson, L., Shaw, J.D., Sparrow, M.D., Summerhayes, C., Terauds, A. & Wall, D.H. 2012. Challenges to the future conservation of the Antarctic. Science 337: 158-159.

Parker, G.C., Rexer-Huber, K., Parker, K.A., Ryan, P.G., Techow, N.M.S.M. & Cuthbert, R.J. 2012. Morphometric and genetic sexing in the Gough Moorhen Gallinula comeri. Ringing and Migration 27: 20-25.

Rexer-Huber, K., Parker, G.C., Ryan, P.G. & Cuthbert, R.J. 2012. Ringing injuries in the Gough Moorhen Gallinula comeri. Ringing & Migration 27: 53-56.

Wanless, R.W., Ratcliffe, N., Angel, A., Bowie, B.C., Cita, K., Hilton, G.M., Kritzinger, P., Ryan P.G. & Slabber, M. 2012. Predation by of Atlantic Petrel chicks by house mice on Gough Island. Animal Conservation 15: 472-479.

Ryan, P.G., Glass, N. & Ronconi, R.A. 2011. The plants and birds of Stoltenhoff and Middle Islands, Tristan da Cunha. Polar Record 47: 86-89.

Cuthbert, R.J., Visser, P., Louw, H. & Ryan, P.G. 2011. Palatability and efficacy of rodent baits for eradicating house mice (Mus musculus) from Gough Island, Tristan da Cunha. Wildlife Research 38: 196-203.

Ryan, P.G. & Ronconi, R.A. 2010. The Tristan Thrush Nesocichla eremita as seabird predator. Ardea 98: 241-243.

Jones, M.G.W. & Ryan, P.G. 2010. Evidence of mouse attacks on albatross chicks on sub-Antarctic Marion Island. Antarctic Science 22: 39-42.

Wanless, R.M., Cooper, J., Slabber, M.J. & Ryan, P.G. 2010. Risk assessment of birds foraging terrestrially at Marion and Gough Islands to primary and secondary poisoning by rodenticides. Wildlife Research 37: 524-530.

Wanless, R.M., Angel, A., Cuthbert, R.J., Hilton, G.M. & Ryan, P.G. 2007. Can predation by invasive mice drive seabird extinctions? Biology Letters 3: 241-244.

Wanless, R.M., Ryan, P.G., Altwegg, R., Angel, A., Cooper, J., Cuthbert, R. & Hilton, G.M. 2009. From both sides: dire demographic consequences of carnivorous mice and longlining for the Critically Endangered Tristan albatrosses on Gough Island. Biological Conservation 142: 1710-1718.

Ryan, P.G. 2008. Taxonomic and conservation implications of ecological speciation in Nesospiza buntings at Tristan da Cunha. Bird Conservation International 18: 30–39.

Ryan, P.G. & Cuthbert, R.J. 2008. The biology and conservation status of the Gough Bunting Rowettia goughensis. Bulletin of the British Ornithologists’ Club 128: 242-253.

Wanless, R.M., Parkes, J., Cooper, J., Fisher, P., Slabber, M. & Ryan, P.G. 2008. An investigation of the susceptibility of mice to poison bait on Gough Island. Wildlife Research 35: 806-811.

Ryan, P.G. & Moloney, C.L. 2002. Breeding behaviour, clutch size and egg dimensions of Nesospiza buntings at Inaccessible Island, Tristan da Cunha. Ostrich 73: 52-58.

Ryan, P.G., Smith, V.R. & Gremmen, N.J.M. 2003. The distribution and spread of alien vascular plants on Prince Edward Island. African Journal of Marine Science 25: 555-561.

Jones, A.G., Chown, S.L., Ryan, P.G., Gremmen, N.J.M. & Gaston, K.J. 2003. A review of conservation threats on Gough Island: a case study for terrestrial conservation in the Southern Oceans. Biological Conservation 113: 75-87.

Ryan, P.G., Cooper, J. & Glass, J.P. 2001. Population status, breeding biology and conservation of the Tristan Albatross Diomedea [exulans] dabbenena. Bird Conservation International 11: 33-46.

Cooper, J. & Ryan, P.G. 2001. The Agreement on the Conservation of Albatrosses and Petrels: implications for research and monitoring at the Prince Edward Islands. South African Journal of Science 97: 78-79.

Ryan, P.G. & Moloney, C.L. 2000. The status of Spectacled Petrels Procellaria conspicillata and other seabirds at Inaccessible Island. Marine Ornithology 28: 93-100.

Huyser, O.A.W., Ryan, P.G. & Cooper, J. 2000. Changes in population size, habitat use and breeding biology of Lesser Sheathbills at Marion Island: impacts of cats, mice and climate change? Biological Conservation 92: 299-310.

Fraser, M.W., Ryan, P.G., Dean, W.R.J., Briggs, D.J. & Moloney, C.L. 1994. Biology of the Tristan Thrush Nesocichla eremita. Ostrich 65: 14-25.

Dean, W.R.J., Milton, S.J., Ryan, P.G. & Moloney, C.L. 1994. The role of disturbance in the establishment of indigenous and alien plants at Inaccessible and Nightingale Islands in the South Atlantic Ocean. Vegetatio 113: 13-24.

Ryan, P.G. 1993. The ecological consequences of an exceptional rainfall event at Gough Island. South African Journal of Science 89: 309-311.

Cooper, J. & Ryan, P.G. 1993. Benign research on a South Atlantic jewel: towards a management plan for Gough Island. George Wright Forum 9 (3-4): 101-112.

Milton, S.J., Ryan, P.G., Moloney, C.L., Cooper, J., Dean, W.R.J. & Medeiros, A.C. Jr. 1993. Disturbance and demography of Phylica arborea (Rhamnaceae) on the Tristan-Gough group of islands. Botanical Journal of the Linnaean Society 111: 55-70.

Roux, J.P., Ryan. P.G., Milton, S.J. & Moloney, C.L. 1992. Vegetation and checklist of Inaccessible Island, central South Atlantic Ocean, with notes on Nightingale Island. Bothalia 22: 93-109.

Ryan, P.G., Moloney, C.L. & Watkins, B.P. 1989. Concern about the adverse effect of introduced mice on island tree Phylica arborea regeneration. South African Journal of Science 85: 626-627.

Ryan, P.G., Watkins, B.P. & Siegfried, W.R. 1989. Morphometrics, metabolic rate and body temperature of the smallest flightless bird: the Inaccessible Island Rail. Condor 91: 465-467.

Avian systematics and evolution (24 papers)

Robertson, B., Stephenson, B., Ronconi, R., Goldstien, S., Shepherd, L., Tennyson, A., Carlile, N. & Ryan, P.G. in press. Phylogenetic affinities of the Fregetta storm-petrels are not black and white. Molecular Phylogenetics Evolution

Moodley, Y., Masello, J., Gopi, M., Cole, T., Thali, M., Alderman, R., Cuthbert, R., Marin, M., Massaro, M., Navarro, J., Phillips, R., Ryan, P.G., Suazo, C., Cherel, Y., Weimerskirch, H. & Quillfeldt, P. 2015. Evolutionary factors affecting the cross-species utility of newly developed microsatellite markers in seabirds. Molecular Ecology Resources doi: 10.1111/1755-0998.12372

Silva, M.C., Matias, R., Wanless, R.M., Ryan, P.G., Stephenson, B.M., Bolton, M., Ferrand, N. & Coelho, M. 2015. Understanding the mechanisms of anti-tropical divergence in the seabird White-faced Storm-petrel (Procellariiformes: Pelagodroma marina) using a multi-locus approach. Molecular Ecology (doi:10.1111/mec.13212)

Ryan, P.G., Klicka, L.B., Barker, F.K. & Burns, K.J. 2013. The origin of finches on Tristan da Cunha and Gough Island, central South Atlantic Ocean. Molecular Phylogenetics & Evolution 69: 299-305.

Ryan, P.G., Bourgeois, K., Dromzeé, S. & Dilley, B.J. 2014. The occurrence of two bill morphs of prions Pachyptila vittata on Gough Island. Polar Biology 37: 727-735.

Alstrom, P., Barnes, K.N., Olsson, U., Barker, F.K., Bloomer, P., Kahn, A.A., Qureshi, M.A., Guillaumet, A., Crochet, P.-A. & Ryan, P.G. 2013. Multilocus phylogeny of the avian family Alaudidae (larks) reveals complex morphological evolution, non-monophyletic genera and hidden species diversity. Molecular Phylogenetics & Evolution 69: 1043-1056.

de Hierro, M.D.G.L., Meléon, M. & Ryan, P.G. 2013. Is carrying feathers a sexually selected trait in House Sparrows? Ethology 119: 1-13.

Jansen van Rensberg, A., Bloomer, P., Ryan, P.G. & Hansson, B. 2012. Genetic variation at the major histocompatibility complex (MHCII beta) in Nesospiza buntings: Effects of genetic drift and shared ancestral polymorphism. BMC Evolutionary Biology 12: 143.

Ryan, P.G., Nupen, L., Rose, B. & Sulieman, A.S. 2010. Geographic variation in Socotra Sparrows Passer insularis. Bulletin of the British Ornithologists’ Club 130: 75-82.

Techow, N.M.S.M., O’Ryan, C., Phillips, R.A., Gales, R., Marin, M., Patterson-Fraser, D., Quintana, F., Ritz, M.S., Thompson, D.R., Wanless, R.M., Weimerskirch, H. & Ryan, P.G. 2010. Speciation and phylogeography of giant petrels Macronectes. Molecular Phylogenetics & Evolution 54: 472–487.

Techow, N.M.S.M., Ryan, P.G. & O’Ryan, C. 2009. Phylogeography and taxonomy of White-chinned and Spectacled Petrels. Molecular Phylogenetics & Evolution 52: 25-33.

de Hierro, M.D.G.L. & Ryan, P.G. 2008. Nest defence and egg rejection in the House Sparrow (Passer domesticus) as protection against conspecific brood parasitism. Behaviour 145: 949-964.

Ritz, M.S., Millar, C., Miller, G.D., Phillips, R.A., Ryan, P.G., Sternkopf, V., Liebers-Helbig, D., Peter, H-U. 2008. Phylogeography of the southern skua complex - rapid colonisation of the Southern Hemisphere during a glacial period and reticulate evolution. Molecular Phylogenetics & Evolution 49: 292-303.

Ryan, P.G., Bloomer, P., Moloney, C.L., Grant, T. & Delport, W. 2007. Ecological speciation in South Atlantic island finches. Science 315: 1420-1423.

Delport, W., Grant, T.J., Ryan, P.G. & Bloomer, P. 2006. Isolation of novel microsatellites, and optimization of published Vidua loci, for evolutionary research on Nesospiza buntings. Molecular Ecology Notes 6: 1180-1183.

Beresford, P., Barker, F.K., Ryan, P.G. & Crowe, T.M. 2005. African endemics span the tree of songbirds (Passeri): Molecular systematics of several evolutionary ‘enigmas’. Proceedings of the Royal Society, London B 272: 849-858.

Ryan, P.G., Wright, D., Oatley, G., Wakeling, J., Cohen, C., Nowell, T.L., Bowie, R.C.K., Ward, V. and Crowe, T.M. 2004. Systematics of Serinus canaries and the status of Cape and Yellow-crowned Canaries inferred from mtDNA. Ostrich 75: 288-294.

Ryan, P.G. 2001. Morphological heritability among a hybrid bunting complex: Nesospiza at Inaccessible Island. Condor 103: 429-438.

Ryan, P.G. 1998. The taxonomic and conservation status of the Spectacled Petrel Procellaria conspicillata. Bird Conservation International 8: 223-235.

Ryan, P.G. & Bloomer, P. 1999. The Long-billed Lark complex: a species mosaic in south-western Africa. Auk 116: 194-208.

Ryan, P.G., Hood, I., Bloomer, P., Komen, J. & Crowe, T.M. 1998. Barlow’s Lark: a new species in the Karoo Lark complex of southwest Africa. Ibis 140: 605-619.

Ryan, P.G. & Bloomer, P. 1997. Geographic variation in Red Lark Certhilauda burra plumage, morphology, song and mitochondrial DNA haplotypes. Ostrich 68: 31-36.

Ryan, P.G., Komen, J. & Moloney, C.L. 1996. The land birds of the Sperrgebiet. Lanioturdus 29: 8-25.

Ryan, P.G., Moloney, C.L. & Hudon, J. 1994. Colour variation and hybridization among Nesospiza buntings on Inaccessible Island, Tristan da Cunha. Auk 111: 314-327.

Collision mortality (8 papers)

Shaw, J.M. & Ryan, P.G. 2015. Stable isotopes reveal regional movement patterns in an endangered bustard. Austral Ecology 40: 198-205. (doi: 10.1111/aec.12194)

Shaw, J.M., Jenkins, A.R., Allan, D.G. & Ryan, P.G. 2015. Population size and trends of Ludwig’s Bustard Neotis ludwigii and other large terrestrial birds in the Karoo, South Africa. Bird Conservation International doi: 10.1017/S0959270914000458.

Shaw, J.M., van der Merwe, R., van der Merwe, E. & Ryan, P.G. 2015. Winter scavenging rates under power lines in the Karoo, South Africa. South African Journal of Wildlife Research 45: 122-126.

Schutgens, M., Shaw, J.M. & Ryan, P.G. 2014. Collision mortality of large birds on power lines: estimating scavenger and search bias in the Karoo, South Africa. Ostrich 85: 39-45.

Shaw, J.M., Dilley, B.J., Davies, D. & Ryan, P.G. 2014. Trapping and capture myopathy in Ludwig’s Bustard. South African Journal of Wildlife Research 44: 16-23.

Jenkins, A.R., Shaw, J.M., Smallie, J.J., Gibbons, B., Visagie, R. & Ryan, P.G. 2011. Power line collisions in Ludwig’s Bustard Neotis ludwigii: measuring rates and modelling demographic effects in the Karoo, South Africa. Bird Conservation International 21: 303-310.

Shaw, J.M., Jenkins, A.R., Smallie, J.J. & Ryan, P.G. 2010. Modelling power line collision risk for the Blue Crane Anthropoides paradiseus in South Africa. Ibis 152: 590-599.

Shaw, J.M., Jenkins, A.R., Ryan, P.G. & Smallie, J.J. 2010. A preliminary survey of avian mortality on power lines in the Overberg, South Africa. Ostrich 81: 109-113.

Shorebird biology and conservation (13 papers)

Paijmans, D.M., Louwenthal, D., Ryan, P.G. & Hockey, P.A.R. 2015. Temporal and spatial differences in three-egg clutch frequency of the African Black Oystercatcher. Ostrich 86: 35-41.

Ryan, P.G. 2013. Medium-term changes in coastal bird communities in the Western Cape, South Africa. Austral Ecology 38: 251-259.

Ryan, P.G. & Visagie, J. 2008. African Black Oystercatchers feeding in a terrestrial habitat. Ostrich 79: 243.

Ryan, P.G. & Sinclair, I. 2006. Mussulo – an important shorebird wintering site in Angola. Wader Study Group Bulletin 109: 120.

Ryan, P.G., Underhill, L.G., Cooper, J. & Waltner, M. 1988. Waders (Charadrii) and other water birds on the coast, adjacent wetlands and offshore islands of the southwestern Cape Province, South Africa. Bontebok 6: 1-19.

Ryan, P.G., Bosman, A.L. & Hockey, P.A.R. 1988. Notes on the foraging behaviour of Flying and Magellanic Flightless Steamer-ducks. Wildfowl 39: 29-33.

Siegfried, W.R., Hockey, P.A.R., Ryan, P.G. & Bosman, A.L. 1988. Sex and plumage-type ratios of the Lesser Magellan Goose in southern Chile. Wildfowl 39: 15-21.

Hockey, P.A.R., Bosman, A.L. & Ryan, P.G. 1987. The maintenance of polymorphism and cryptic mimesis in the limpet Scurria veriabilis by two species of Cinclodes (Aves: Furnariinae) in central Chile. Veliger 30: 5-10.

Ryan, P.G., Cooper, J., Hockey, P.A.R. & Berruti, A. 1986. Waders (Charadrii) and other water birds on the coast and adjacent wetlands of Natal, 1980-81. Lammergeyer 36: 1-33.

Ryan, P.G. & Cooper, J. 1985. Waders (Charadrii) and other coastal birds of the northwestern Cape Province, South Africa. Bontebok 4: 1-8.

Hockey, P.A.R. & Ryan, P.G. 1985. Waders on rocky shores: does resource depletion lead to competition? South African Journal of Science 81: 698.

Ryan, P.G., Cooper, J. & Stutterheim, C.J. 1984. Waders (Charadrii) and other coastal birds of the Skeleton Coast, South West Africa. Madoqua 14: 71-78.

Ryan, P.G., Cooper, J., Stutterheim, C.J. & Loutit, R. 1984. An annotated list of the birds of the Skeleton Coast Park. Madoqua 14: 79-90.

Other conservation papers (12 papers)

Maphisa, D.H., Donald, P.F., Buchanan, G.M. & Ryan, P.G. 2009. Habitat use, distribution and breeding ecology of the globally threatened Rudd’s Lark and Botha’s Lark in eastern South Africa. Ostrich 80: 19-28.

Lauret-Stepler, M., Bourjea, J., Roos, D., Pelletier, D., Ryan, P.G., Ciccione, S. & Grizel, H. 2007. Reproductive seasonality and trend of Chelonia mydas in the SW Indian Ocean: a 20 year study based on track counts. Endangered Species Research 3: 217-227.

Owino, A.O. & Ryan, P.G. 2007. Recent papyrus swamp habitat loss and conservation implications in western Kenya. Wetlands Ecology & Management 15: 1-12.

Muntifering, J.R., Dickman, A.J., Perlow, L.M., Hruska, T., Ryan, P.G., Marker, L.L. & Jeo, R.M. 2006. Managing the matrix for large carnivores: a novel approach and perspective from cheetah (Acinonyx jubatus) habitat suitability modelling. Animal Conservation 9: 103-112.

Owino, A.O. & Ryan, P.G. 2006. Habitat associations of papyrus specialist birds at three papyrus swamps in western Kenya. African Journal of Ecology 44: 438-443.

Ryan, P.G., Sinclair, I., Cohen, C., Mills, M.S.L., Spottiswoode, C.N. & Cassidy, R. 2004. The conservation status and vocalisations of threatened birds from the scarp forest of Cuanza Sul, Angola. Bird Conservation International 14: 247-260.

Ryan, P.G. & Spottiswoode, C.N. 2003. Long-billed Tailorbirds (Orthotomus moreaui) rediscovered at Serra Jeci, northern Mozambique. Ostrich 74: 141-145.

Ndang'ang'a, P.K., du Plessis, M.A., Ryan, P.G. & Bennun, L.A. 2002. Grassland decline in Kinangop Plateau, Kenya: implications for conservation of Sharpe's Longclaw (Macronyx sharpei). Biological Conservation 107: 341-350.

Ryan, P.G., Bento, C., Cohen, C., Graham, J., Parker V. & Spottiswoode, C. 1999. The avifauna and conservation status of the Namuli Massif, northern Mozambique. Bird Conservation International 9: 315-331.

Joubert, D.F. & Ryan, P.G. 1999. Differences in mammal and bird assemblages between commercial and communal range-lands in the Succulent Karoo, South Africa. Journal of Arid Environments 43: 287-299.

Ryan, P.G., Cowling, R.M. & Costanza, R. 1995. Valuing fynbos ecosystems: is the fynbos worth conserving? South African Journal of Science 91: 572-573.

Ryan, P.G. & Hockey, P.A.R. 1995. Is the Hottentot Buttonquail really endangered? Ostrich 66: 92-93.

Sundry papers (23 papers)

Zoghby, B., Little, R.M., Ryan, P.G. & Hockey, P.A.R. in press. Patterns of roost site selection and use by Southern Ground-Hornbills in the South African lowveld. Ostrich

Zoghby, B., Ryan, P.G., Little, R.M., Reid, T. & Hockey, P.A.R. 2015. Seasonal changes in movement and habitat use by Southern Ground-Hornbills in the South African lowveld. Ostrich 89: 87-95. (doi: 10.2989/00306525.2015.1030794).

Bester, M.N., Ryan, P.G., Bester, W.A. & Glass, T. 2014. Vagrant Antarctic fur seals at the Tristan da Cunha Islands. Polar Biology 37: 1701-1703.

Ryan, P.G. 2013. The effect of wind direction on flying fish counts. African Journal of Marine Science 35: 585-587.

Treasure, A.M., Moloney, C.L., Bester, M.N., McQuaid, C.D., Findlay, K.P., Best, P.B., Cowan, D.A., de Bruyn, P.J.N., Dorrington, R.A., Fagereng, A., Froneman, P.W., Grantham, G.H., Hunt, B.P.V., Meiklejohn, K.I., Pakhomov, E.A., Roychoudhury, A.N., Ryan, P.G., Smith, V.R., Chown, S.L. and Ansorge, I.J. 2013. South African research in the Southern Ocean: new opportunities and challenges. South African Journal of Science doi: 10.1590/sajs.2013/a009

Ryan, P.G. & Branch, G.M. 2012. The November 2011 irruption of buoy barnacles Dosima fascicularis in the Western Cape, South Africa. African Journal of Marine Science 35: 157-162.

Crous, P.W., Groenewald, J.Z., Ryan, P.G. & Wingfield, M.J. 2012. Fungal Planet description sheet 147: Seiridium phylicae. Persoonia 29: 186-187.

Cooper, J., Bester, M.N., Chown, S.L., Crawford, R.J.M., Daly, R., Heyns, E., Lamont, T., Ryan, P.G. & Shaw, J.D. 2009. Biological survey of the Prince Edward Islands, December 2008. South African Journal of Science 105: 317-320.

Ryan, P.G., Porter, R.F., Rose, B. & Sulieman, A.S. 2009. Juvenile plumage of the Socotra Bunting Emberiza socotrana. Sandgrouse 31: 109-111.

Best, P.B., Glass, J.P., Ryan, P.G. & Dalebout, M. 2009. Cetacean records from Tristan da Cunha, South Atlantic. Journal of the Marine Biology Association UK 89: 1023-1032.

Bester, M.N., Visagie, J. & Ryan, P.G. 2009. Summer survey of fur seals at Prince Edward Island, Southern Ocean. African Journal of Marine Science 31: 451-455.

Ansorge, I.J., Roman, R., Durgadoo, J.V., Ryan, P.G., Dlamini, L., Gebhardt, Z., Rainier, S., Smith, M., Mtonsti, T. & Lutjeharms, J.R.E. 2008. The first oceanographic survey of the Conrad Rise. South African Journal of Science 104: 333-336.

Ryan, P.G. & Marshall, H.K. 2005. Variation in the wing-clapping display of Clapper Larks. Ostrich 76: 73-77.

Bragg, C.J., Donaldson, J.D. & Ryan, P.G. 2005. Cape porcupines as ecosystem engineers: aspects of density and disturbance in a geophyte-rich semi-arid environment. Journal of Arid Environments 61: 261-275.

Ryan, P.G. 2004. Nest sites and clutch sizes of House Sparrows at Dassen Island, South Africa. Ostrich 75: 165-167.

Bester, M.N., Ryan, P.G. & Dyer, B.M. 2003. Population numbers of fur seals at Prince Edward Island, Southern Ocean African Journal of Marine Science 25: 549-554.

Crawford, R.J.M., Underhill, L.G., Bester, M.N., Cooper, J., Greyling, M.D., Meÿer, M.A., Petersen, S.L., Ryan, P.G. & Wilkinson, W. 2003. The Prince Edward Islands Millennium Expedition. South African Journal of Science 99: 407-409.

Branch, W.R. & Ryan, P.G. 2001. Additions to the Mozambique herpetofauna: two new lizards from the Namuli Massif, Mozambique. Herpetological Review 32: 281-282.

Cooper, J., Siegfried, W.R., Ryan, P.G., Crafford, J.E. & Stock, W.D. 1991. Effects of ornithogenic products on ecosystem structure and functioning: a new South African Biological Antarctic Research Programme. South African Journal of Science. 87: 223-226.

Cockroft, V.H., Peddemors, V.M., Ryan, P.G. & Lutjeharms, J.R.E. 1990. Cetaceans associated with Agulhas Current rings in the Southern Ocean. South African Journal of Antarctic Research 20: 64-67.

Dastych, H., Ryan, P.G. & Watkins, B.P. 1990. Notes on Tardigrada from western Dronning Maud Land (Antarctica), with a description of two new species. Entomological Mitteilungen Zoological Museum Hamburg Bd. 10: 57-66.

Ryan, P.G. & Watkins, B.P. 1989. The influence of physical factors and ornithogenic products on plant and arthropod abundance at an inland nunatak group in Antarctica. Polar Biology 10: 151-160.

Ryan, P.G., Watkins, B.P., Lewis Smith, R.I., Dastych, H., Eicker, A., Foissner, W., Heatwole, H., Miller, W.R. & Thompson, G. 1989. Biological survey of Robertskollen, western Dronning Maud Land: area description and preliminary species list. South African Journal of Antarctic Research 19: 10-20.

Books (10):

Sinclair, I., Hockey, P., Tarboton, W. & Ryan, P. 2011. SASOL Birds of Southern Africa (Fourth edition). Struik Nature, Cape Town. 464 pp. (ISBN 978 1 77007 925 0). Also in Afrikaans (ISBN 978 1 77007 926 7)

Sinclair, I. & Ryan, P.G. 2010. Birds of Africa South of the Sahara. (Second edition). Struik Nature, Cape Town. 768 pp. (ISBN 978 1 77007 623 5)

Terauds, A., Cooper, J., Chown, S.L. & Ryan, P.G. 2010. Marion and Prince Edward: Africa’s southern islands. SunMedia, Stellenbosch. 176 pp. (ISBN 978 1 920338 42 8).

Sinclair, I. & Ryan, P.G. 2009. Complete Photographic Field Guide: Birds of Southern Africa. Struik Nature, Cape Town. 432 pp. (ISBN 978 1 77007 388 3). Also in Afrikaans (ISBN 978 1 77007 589 4)

Ryan, P.G. 2008. The birds and other wildlife of Kirstenbosch. Struik, Cape Town. 56 pp. (ISBN 978 1 77007 725 6)

Ryan, P.G. (ed.) 2007. Field guide to the animals and plants of Tristan da Cunha and Gough Island. Pisces Publications, Newbury. 162 pp. (ISBN 1 874357 33 1)

Hockey, P.A.R., Dean, W.R.J. & Ryan, P.G. (eds). 2005. Roberts’ Birds of Southern Africa (Seventh edition). The Trustees of the John Voelcker Bird Book Fund, Cape Town. 1296 pp. (ISBN 0 620 34053 3)

Sinclair, I. & Ryan, P.G. 2003. Birds of Africa south of the Sahara. Struik, Cape Town. 760 pp. (ISBN 1 86872 857 9)

Ryan, P.G. 2001. Practical birding. Struik, Cape Town. 96 pp. (ISBN 1 86872 608 8)

Hockey, P.A.R., Underhill, L.G., Neatherway, M. & Ryan, P.G. 1989. Atlas of the birds of the southwestern Cape. Cape Bird Club, Cape Town, 236 pp. (ISBN 0 620 14461 0)

Book chapters, management plans and conference proceedings/abstracts (44):

Ryan, P.G. 2015. A brief history of marine litter research. In: Bergmann, M., Gutow, L. & Klages, M. (eds) Marine Anthropogenic Litter. Springer International Publishing, Cham. Pp. 1-25.

GESAMP (2015). “Sources, fate and effects of microplastics in the marine environment: a global assessment” (Kershaw, P. J., ed.). (IMO/FAO/UNESCO-IOC/UNIDO/WMO/ IAEA/UN/UNEP/UNDP Joint Group of Experts on the Scientific Aspects of Marine Environmental Protection). Rep. Stud. GESAMP No. 90, 96 pp. (contributors: Anderson, A., Andrady, A., Arthur, C., Baker, J., Bouwman, H., Gall, S., Hidalgo-Ruz, V., Köhler, A., Law, K.L., Leslie, H., Kershaw, P., Pahl, S., Potemra, J., Ryan, P., Joon Shim, W., Thompson, R., Takada, H., Turra, A., Vethaak, D. & Wyles, K.)

Shaw, J. & Ryan, P. 2012. Keeping an eye on bird routes. In: Clavaud, R. (ed). Sustainable development in Africa and satellites. Suds-Concepts, France. Pp. 76-77.

Cooper, J., Cuthbert, R.J., Gremmen, N.J.M., Ryan, P.G. & Shaw, J.D. 2011. Earth, fire and water: applying novel techniques to eradicate the invasive plant, procumbent pearlwort Sagina procumbens, on Gough Island, a World Heritage Site in the South Atlantic. In: Veitch, C.R., Clout, M.N. & Towns, D.R. (eds). Island invasives: Eradication and management. IUCN, (International Union for Conservation of Nature), Gland, Switzerland. Pp. 162-165 (ISBN 978 2 8317 1291 8).

Cuthbert, R., Visser, P., Louw, H., Rexer-Huber, K., Parker, G. & Ryan, P. 2011. Preparations for the eradication of mice from Gough Island: results of bait acceptance trials above ground and around cave systems. In: Veitch, C.R., Clout, M.N. & Towns, D.R. (eds). Island invasives: Eradication and management. IUCN, (International Union for Conservation of Nature), Gland, Switzerland. Pp. 47-50 (ISBN 978 2 8317 1291 8).

Jaramillo, A., Rising, J.D., Copete, J.L., Ryan, P.G. & Madge, S.C. 2011. Species accounts of Emberizidae. In: del Hoyo, J., Elliott, A. & Christie, D. (eds) Handbook of the Birds of the World, Vol. 16. Lynx, Barcelona. Pp. 502-683. (ISBN 978 84 96553 78 1).

Ryan, P.G. 2009. Tristan da Cunha and Gough Island. In R. Gillespie & D. Clague (eds) Encyclopedia of Islands. University of California Press, Berkeley. Pp. 929-932. (ISBN 978 0 520 25649 1).

Ryan, P.G. & Bester, M.N. 2008. Pelagic predators. In: Chown, S.N. & Froneman, W. (eds) The Prince Edwards Archipelago: Land-sea interactions in a changing ecosystem. Sun Media, Stellenbosch. Pp. 121-164. (ISBN 978-1-920109-85-1).

Ryan, P.G. 2006. Inaccessible Island Seabird Monitoring Manual. RSPB Research Report 16: 1-32. (ISBN 1 901930 70X).

Ryan, P.G. 2006. Family Cisticolidae. In: del Hoyo, J., Elliott, A. & Christie, D. (eds) Handbook of the Birds of the World, Vol. 11. Lynx, Barcelona. Pp. 378-419. (ISBN 978 96553 06 4).

Ryan, P.G., Dean, W.R.J., Madge, S. & Pearson, D. 2006. Species accounts of Cisticolidae. In: del Hoyo, J., Elliott, A. & Christie, D. (eds) Handbook of the Birds of the World, Vol. 11. Lynx, Barcelona. Pp. 420-490. (ISBN 978 96553 06 4).

Ryan, P.G., de Juana, E., Suárez, F., Alström, P. & Donald, P.F. 2004. Species accounts of Alaudidae. In: del Hoyo, J., Elliott, A. & Christie, D. (eds) Handbook of the Birds of the World, Vol. 9. Lynx, Barcelona. Pp. 542-601. (ISBN 978 87334 69 5).

de Juana, E., Suárez, F. & Ryan, P.G. 2004. Family Alaudidae (Larks). In: del Hoyo, J., Elliott, A. & Christie, D. (eds) Handbook of the Birds of the World, Vol. 9. Lynx, Barcelona. Pp. 496-541. (ISBN 978 87334 69 5).

Tuck, G.N., Polacheck, T., Croxall, J.P., Weimerskirch, H., Ryan, P.G., Nel, D.C., Wayte, S. & Bulman, C.M. 2004. Modelling the impact of fishery incidental mortality on three populations of Wandering Albatross. In: Tuck, G.N. (ed.). A comprehensive study of the ecological impacts of the worldwide pelagic longline industry: southern hemisphere studies. CSIRO Marine Research, Hobart. Pp. 149-223.

Glass, J.P. & Ryan, P.G. 2003. Conservation challenges in small communities: conservation management in the Tristan islands. In: Pienkowski, M. (ed.) A sense of direction: a conference on conservation in UK Overseas Territories and other small island communities. UK Overseas Territories Conservation Forum, www.ukotcf.org. Pp. 139-147.

Ryan, P.G. & Glass, J.P. 2001. Inaccessible Island Nature Reserve Management Plan. Government of Tristan da Cunha, Edinburgh, Tristan da Cunha. 65 pp.

Ryan, P.G., Purves, M. & Cooper, J. 2001. Problems with pirates: toothfish longlining and seabird bycatch at the Subantarctic Prince Edward Islands. In: Melvin, E.F. & Parrish, J.K. (eds) Seabird bycatch: trends, roadblocks, and solutions. University of Alaska Sea Grant, Fairbanks. Pp. 195.

Bloomer, P., Ryan, P.G. & Crowe, T.M. 1998. Some surprises from a molecular phylogeny for southern African larks. Ostrich 69: 397 [International Ornithological Congress abstract]

Nel, D.C., Hurford, J. & Ryan, P.G. 1998. Why is there sex-biased mortality of seabirds in the Prince Edward Island toothfish fishery? Ostrich 69: 442 [International Ornithological Congress abstract]

Ryan, P.G. & Marshall, H. 1998. The clapping arms race: why do some Clapper Larks not bother to play the game? Ostrich 69: 266 [International Ornithological Congress abstract]

Turpie, J.K. & Ryan, P.G. 1998. The value of birds and birding in South Africa. Ostrich 69: 455-456 [International Ornithological Congress abstract]

Ryan, P.G. 1997. Family Diomedeidae: albatrosses. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 13-15, 755-757.

Ryan, P.G. 1997. Family Procellariidae: petrels, shearwaters and prions. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 16-20, 757-761.

Ryan, P.G. 1997. Family Oceanitidae: storm petrels. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 21-23, 761-762.

Ryan, P.G. 1997. Family Phaethontidae: tropicbirds. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 762.

Ryan, P.G. 1997. Family Fregatidae: frigatebirds. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 763.

Ryan, P.G. 1997. Family Chionididae: sheathbills. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 772.

Ryan, P.G. & Crawford, R.J.M. 1997. Family Sulidae: gannets and boobies. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 28-29, 762-763.

Ryan, P.G., Crawford, R.J.M. & Whittington, P.A. 1997. Family Spheniscidae: penguins. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 4-5, 755.

Ryan, P.G., Williams, A.J., Crawford, R.J.M., Underhill, L.G., Brooke, R.K., Herremans, M. & Simmons, R.E. 1997. Family Laridae: skuas, gulls and terns. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 1: Non-passerines. BirdLife South Africa, Johannesburg. Pp. 459-484, 772-775.

Dean, W.R.J., Allan, D.G., Herremans, M., Ryan, P.G. & Cohen, C. 1997. Family Alaudidae: larks. In: Harrison, J.A., Allan, D.G., Underhill, L.G., Herremans, M., Tree, A.J., Parker, V. & Brown, C.J. (eds) The atlas of southern African birds. Vol. 2: Passerines. BirdLife South Africa, Johannesburg. Pp. 4-47, 687.

Gregory, M.R. & Ryan, P.G. 1997. Pelagic plastics and other seaborne persistent synthetic debris: a review of Southern Hemisphere perspectives. In: Coe, J.M. & Rogers, D.B. (eds) Marine debris: sources, impacts and solutions. Springer-Verlag, New York. Pp. 49-66.

Ryan, P.G. 1996. Identifying problem species: the Karoo Lark complex (Karoo, Red, Dune and Barlow’s Larks). In: Sinclair, J.C. & Hockey, P.A.R. The larger illustrated guide to birds of southern Africa. Struik, Cape Town. Pp. 423-424.

Allan, D.G. & Ryan, P.G. 1996. Morphometrics, sex ratio, molt, and stomach contents of Blue Cranes in the western Cape Province, South Africa. In: Beilfuss, R.D., Tarboten, W.R. & Sichuki, N.N. (eds) Proc. 1993 Afr. Crane & Wetland Training Workshop. International Crane Foundation, Baraboo, Wisconsin. Pp. 377-387.

Ryan, P.G., Hood, I., Bloomer, P. & Komen, J. 1995. Barlow’s Lark - a near endemic to the Sperrgebiet. Pp. 54-55 in Pallett, J. (ed.) The Sperrgebiet: Namibia’s least known wilderness. NAMDEB, Windhoek.

Cooper, J. & Ryan, P.G. 1995. The conservation status of Gough Island. In: Dingwall, P.R. (ed.) Progress in the conservation of the Subantarctic Islands. World Conservation Union: Conservation of the Southern Polar Regions 2: 71-84.

Cooper, J., Ryan, P.G. & Andrew, T.G. 1995. The conservation status of the Tristan da Cunha Islands. In: Dingwall, P.R. (ed.) Progress in the conservation of the Subantarctic Islands. World Conservation Union: Conservation of the Southern Polar Regions 2: 59-70.

Moloney, C.L & Ryan, P.G. 1995. Antarctic marine foodwebs. In: Nierenberg, W.A. (ed.) Encyclopedia of Environmental Biology, Vol. 1. Academic Press, San Diego. Pp 53-69.

Ryan, P.G. & Siegfried, W.R. 1994. The viability of small populations of birds: an empirical investigation of vulnerability. In: Remmert, H. (ed.) Minimum animal populations. Ecological Studies 106: 3-22. Springer-Verlag, Berlin. [reprinted in German and English in Nationalpark Berchtesgaden Forschungsbericht 27: 6-31.]

Cooper, J. & Ryan, P.G. 1994. Management Plan for the Gough Island Wildlife Reserve. Government of Tristan da Cunha, Edinburgh, Tristan da Cunha. 96 pp.

Crowe, T.M., Ryan, P.G., Essop, M.F., Brooke, R.K., Hockey, P.A.R. & Siegfried, W.R. 1994. Species as the 'currency' of conservation: the Karoo/Dune/Red Lark complex of south-western Africa. Pp 229-234 in Forey, P.L., Humphries, C.J. & Vane-Wright, R.I. (eds) Systematics and Conservation Evaluation. Clarendon Press, Oxford.

Ryan, P.G. 1990. The marine plastic debris problem off southern Africa: types of debris, their environmental effects, and control measures. Proc. Second Int. Conf. on Marine Debris, NOAA Tech. Mem., NMFS, SWFC 154: 85-102.

Ryan, P.G. 1990. The effects of ingested plastic and other marine debris on seabirds. Proc. Second Int. Conf. on Marine Debris, NOAA Tech. Mem., NMFS, SWFC 154: 623-634.

Ryan, P.G. & Rose, B. 1989. Migrant seabirds. In: Payne, A.I.L. & Crawford, R.J.M. (Eds). Oceans of life off southern Africa. Vlaeberg, Cape Town, pp. 274-287.