Research Associates

Dr Grant Joseph

Dr Grant Stuart Joseph


Grant is both a practicing medical practitioner and research scientist. Following 10 years of medical practice, he completed an MSc in Conservation Biology and a PhD in Zoology. Research interests include Conservation Biology, Landscape Ecology, Community Ecology, Restoration Ecology, ornithology, and the emerging threats to human health from global environmental change. He is presently investigating the influence of termitaria on biodiversity and resilience in dystrophic savannah systems, restoration of semi-arid systems, the effects of environmental change on bird functional distribution across gradients of aridity, and on ecological systems and human health. He believes that through education and shared knowledge with the general public and commercial institutions, scientists can play a decisive role in achieving a sustainable future in which the wonder of four billion years of life is respected, and better understood by all.

Recent peer-reviewed publications


Dean, W.R.J., Seymour, C.L. , Joseph, G.S. and Foord, S.H. 2019. A Review of the Impacts of Roads on Wildlife in Semi-Arid Regions. Diversity 11(5). doi:10.3390/d11050081.

Joseph, G.S., Muluvhahothe, M.M., Seymour, C.L., Munyai, T.C., Bishop, T.R. and Foord, S. 2019. Stability of Afromontane ant diversity decreases across an elevation gradient. Global Ecology and Conservation.

Seymour, C. and Joseph, G.S. 2019. Ecology of smaller animals associated with savanna woody plants. The value of the finer details.  pg 183-212. In Scogings, P.F. and Sankaran, M. (eds). Savanna woody plants and large herbivores,ISBN: 978-1-119-08110-4. 784 pages.

Seymour, C.L., Milton, S.J., Altwegg, R., Joseph, G.S. and Dean, W.R.J. 2019. Addition of Nitrogen Increases Variability of Vegetation Cover in an Arid System with Unpredictable Rainfall. Ecosystems: 1-13. 


Dean, W.R.J, Seymour, C.L. and Joseph, G.S. 2018. Linear structures in the Karoo, South Africa, and their impacts on biota. African Journal of Range & Forage Science 35 (3-4), 223-232.

Joseph, G.S., Seymour, C.S., Coetzee, B.W.T., Ndlovu, M., Luana Deng, L., Fowler, K., Hagan, J., Brooks, B.J., Seminara, J.A. and Foord, S.H. 2018. Elephants, termites and mound thermoregulation in a progressively warmer world. Landscape Ecology.

Simba, L.D. Foord; S.H., Thebault, E. ; Van Veen, F.J., Joseph, G.S. and Seymour, C.L.. 2018. Indirect interactions between crops and natural vegetation through flower visitors: The importance of temporal as well as spatial spillover . Agriculture, Ecosystems and Environment 253: 148-156.


Joseph, G.S., Mauda, E.V., Seymour, C.L., Thinandavha C. Munyai, T.C., Dippenaar-Schoeman, A. and Foord, S.H. 2017. Landuse change in savannas disproportionally reduces functional diversity of invertebrate predators at the highest trophic levels: spiders as an example. Ecosytems.21(5): 930-942.

Joseph, G.S., Seymour, C.L. and Foord, S.H.. 2017. The effect of infrastructure on the invasion of a generalist predator: Pied crows in southern Africa as a case-study. Biological Conservation (205) 11-15.

Mauda, E.V., Joseph, G.S., Seymour, C.L., Munyai, T.C., Foord, S.H. 2017. Changes in landuse alter ant diversity, assemblage composition and dominant functional groups in African savannas. Biodiversity and Conservation 1-19.

Modiba, R.V, Joseph, G.S., Seymour, C.L., Fouché, P. and Foord, S.H. 2017. Restoration of riparian systems through clearing of invasive plant species improves functional diversity of Odonate assemblages. Biological Conservation 214, 46-54.


Joseph, G.S., Seymour, C.L., Coetzee, B.T.W., Ndlovu, M., de la Torre, A., Suttle, R., Hicks, N., Oxley, S. and Foord, S.H. 2016. Microclimates mitigate against hot temperatures in dryland ecosystems: termite mounds as an example.  Ecosphere 7(11) e01509. doi: 10.1002/ecs2.1509.

Seymour, C.L., Joseph, G.S., Makumbe, M., Cumming, G.S., Mahlangu & Cumming, D.H.M. 2016. Woody species composition in an African savanna: determined by centuries of termite activity but modulated by 50 years of ungulate herbivory. Journal of Vegetation Science Doi: 10.1111/jvs.12393.


Seymour, C.L., Simmons, R.E., Joseph, G.S. & Slingsby, J.A. 2015. On bird functional diversity: species richness and functional differentiation show contrasting responses to rainfall and vegetation structure in an arid landscape. Ecosystems. Doi: 10.1007/s10021-015-9875-8.

Joseph, G.S., Makumbe, M., Seymour, C.L., Cumming, G.S., Mahlangu, Z., and Cumming, D.H.M. 2015. Termite mounds mitigate against 50 years of herbivore-induced reduction of functional diversity of savanna woody plants. Landscape Ecology 30: 2161-2174. doi:10.1007/s10980-015-0238-9.


Joseph, G.S., Seymour, C., Cumming, G.S., Cumming, D.H.M., and Mahlangu, Z. 2014. Termite mounds increase functional diversity of woody plants in African savannas. Ecosystems. DOI: 10.1007/s10021-014-9761-9

Seymour, C.L., Milewski, A.V., Mills, A.J., Joseph, G.S., Cumming, G.S., Cumming, D.H.M. & Mahlangu, Z. 2014. Do the large termite mounds of Macrotermes concentrate micronutrients in addition to macronutrients in nutrient-poor African savannas? Soil Biology and Biochemistry 68:105-105.


Dean, W.R.J., Franke, U., Joseph, G.S., Gonçalves, F.M., Mills, M.S.L., Milton, S.J., Monadjem, A. & Oschadleus, H.D. 2013. Further breeding records for birds (aves) in Angola. Durban Natural Science Museum Novitates (36) 1-10.

Joseph, G.S., Seymour, C., Cumming, G.S., Cumming, D.H.M. & Mahlangu, Z. 2013. Termite mounds as islands: woody plant assemblages relative to termitarium size and soil properties. Journal of Vegetation Science 24(4) 702-711.

Joseph, G.S., Seymour, C., Cumming, G.S., Mahlangu, Z. & Cumming, D.H M. 2013. Escaping the flames: large termitaria as refugia from fire in miombo woodland. Landscape Ecology doi:10.1007/s10980-013-9897-6.

Seymour, C., Milewski, A.V., Mills, A.J., Joseph, G.S., Cumming, G.S., Cumming, D.H.M., and Mahlangu, Z. 2013. Do the large termite mounts of Macrotermes concentrate both micro- and macronutrients in nutrient-poor African savannas? Soil Biology and Biochemistry (68) 105-105.


Dean, W.R.J., Franke, U., Joseph, G.S., Gonçalves,F.M.. Mills, M.S.L., Milton,S.J., Monadjem, A. & Oschadleus, H.D. 2012. Type specimens in the bird collection at Lubango, Angola. Bulletin of the British Ornithologists' Club 132:41-45.


Joseph, G.S., Cumming, G.S., Cumming, D.H.M., Mahlangu, Z., Altwegg, R. & Seymour, C. 2011. Large termitaria act as refugia for tall trees, deadwood and cavity-using birds in a miombo woodland. Landscape Ecology 26:439-448.


Child, M.F., Milton, S.J., Dean, W. R. J., Lipsey, M. K., Puttick, J., Hempson, T. N., Mann, G. K., Babiker, H., Chaudrey, J., Humphrey, G., Joseph, G.S., Okes, N. C., Potts, R. and Wistebaar, T. 2010. Tree-grass coexistence in a flood-disturbed, semi-arid savanna system. Landscape Ecology 25(2):315-326.

Mills, M.S.L., Franke, U. Joseph, G.S., Miato, F., Milton, S., Monadjem, A., Oschadleus, D. & Dean, W.R.J. 2010. Cataloguing the Lubango Bird Skin Collection: towards an atlas of Angolan bird distributions. Bulletin of the African Bird Club 17:43-53.

Seymour, C.L., Milton, S.J., Joseph, G.S., Dean, W.R.J., Ditlhobolo, T. & Cumming, G.S. 2010. Twenty years of rest returns grazing potential, but not palatable plant diversity, to Karoo rangeland, South Africa. Journal of Applied Ecology 47:859-867.